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Computational models of cardiac disease — landscape survey

Provenance/reference document for dismech curation of computational_models: blocks on cardiac disorder entries. Compiled July 2026. All PMIDs below were verified against PubMed records or the publisher page; the four flagged as "search-confirmed" were taken from search-result summaries with matching PubMed links but not individually re-opened — double-check before hard-committing them to a KB entry.

Why this matters for dismech

Cardiac electrophysiology is the most model-rich subfield of physiology, and the dismech schema already supports it: the top-level computational_models: slot (ComputationalModel class) captures ODE/PDE and other in-silico models, with model_type: KINETIC for ODE systems, variables (each ModelVariable with a native dataset_identifier, unit, and ontology mappings_list), and ModelVariableDescriptor thresholds/severity_scale that link a state-variable value to an HP phenotype. Evidence uses evidence_source: COMPUTATIONAL. The prior worked example was CKD-Mineral_Bone_Disorder.yaml (Peterson-Riggs ODE model, BIOMD0000000613); Long_QT_Syndrome.yaml is now the first cardiac worked example (ORd, CiPA IKr-dynamic ORd, ToR-ORd).

Models organized by spatial scale

Cardiac modeling is canonically multiscale: ODEs describe subcellular fluxes and single-cell membrane dynamics; those cell models become the reaction term inside tissue/organ reaction-diffusion PDEs; continuum finite-element mechanics (plus fluid-structure interaction for blood) closes the electromechanical loop.

Subcellular / organelle (ODE systems)

Sub-system What is modeled Representative models
Mitochondrial bioenergetics TCA cycle, oxidative phosphorylation, ATP/ADP, NADH, membrane potential, matrix Ca²⁺ Cortassa–Aon–Marbán–Winslow–O'Rourke 2003; ECME extension (Cortassa 2006)
Ca²⁺ handling / SR L-type Ca²⁺ current, RyR2 Ca²⁺-induced Ca²⁺ release, SERCA uptake, compartmental Ca²⁺ Shannon–Bers 2004 (reference SR/Ca²⁺ framework)
Myofilament / contraction Cross-bridge cycling, Ca²⁺–troponin binding, length/tension dependence Rice et al. 2008; Land et al. (used with ToR-ORd)

These ODE sub-models are usually plugged into a single-cell electrophysiology model rather than run alone.

Single-cell cardiomyocyte electrophysiology (Hodgkin–Huxley lineage)

Systems of ODEs for membrane potential + gating variables + ion concentrations:

  • Hodgkin–Huxley 1952 (squid axon) — mathematical template, not cardiac.
  • Noble 1962 — first cardiac model (Purkinje fibre).
  • Beeler–Reuter 1977 — first mammalian ventricular AP.
  • Luo–Rudy I (1991) / II dynamic (1994) — guinea-pig ventricle; 1990s workhorse.
  • Courtemanche 1998 / Nygren 1998 — foundational human atrial cell models (AF substrate).
  • Shannon–Bers 2004 — rabbit ventricle, gold-standard Ca²⁺ handling.
  • ten Tusscher 2004 / ten Tusscher–Panfilov 2006 — first widely adopted human ventricular models, efficient enough for tissue/organ use.
  • Grandi–Pasqualini–Bers 2010 — human ventricular AP + Ca²⁺ transient; strong for CaMKII / heart-failure remodeling.
  • O'Hara–Rudy "ORd" 2011 — modern reference human ventricular model; CiPA consensus model.
  • ToR-ORd (Tomek et al. 2019) — ORd refinement (corrected reversal potentials, reformulated I_CaL, replaced hERG), validated in disease and drug block, cell-to-ECG.
  • Paci et al. hiPSC-CM models (2013 →) — human iPSC-derived cardiomyocyte models (the main in-vitro human platform).

Key ventricular currents: I_Na, I_CaL, I_Kr (hERG — central to drug safety), I_Ks, I_K1, I_to, I_NaCa, I_NaK, plus SR fluxes and CaMKII regulation.

Tissue / organ (reaction-diffusion PDEs)

  • Monodomain — single reaction-diffusion PDE (cell ionic model = reaction term + diffusion term); cheaper, assumes equal anisotropy ratios.
  • Bidomain — coupled intra/extracellular PDE pair; required for defibrillation, external stimulation, ECG/body-surface potentials, unequal anisotropy.
  • Whole-heart / organ — 3D geometries + fiber orientation for arrhythmia mechanism, ablation planning, in-silico drug trials (reviewed in Clayton 2011).

Mechanics / finite-element and fluid-structure

  • Nonlinear anisotropic hyperelastic constitutive laws (Guccione, Holzapfel–Ogden) + active tension → electromechanical whole-heart models.
  • Fluid-structure interaction couples deforming myocardium/valves to intracavitary blood flow (Navier–Stokes). Basis of "cardiac digital twin" pipelines.

Mapping model types to diseases (and candidate dismech entries)

Disease Typical modeling approach dismech entry
Long QT syndrome Ventricular ODE (ORd/ToR-ORd/Grandi) with I_Kr↓ (LQT2), I_Ks↓ (LQT1), late I_Na↑ (LQT3) → APD↑ → EADs Long_QT_Syndrome ✅ (done)
Drug-induced TdP CiPA IKr-dynamic ORd + multi-channel pharmacology → qNet Long_QT_Syndrome ✅, Torsade_De_Pointes_Syndrome_With_Short_Coupling_Interval
Brugada syndrome Ventricular cell + tissue, reduced I_Na (SCN5A) / altered I_to, transmural heterogeneity Brugada_Syndrome
Short QT syndrome Ventricular ODE with gain-of-function K⁺ currents → APD↓ Short_QT_Syndrome
CPVT Ca²⁺-handling models (Shannon–Bers/Grandi) with RyR2/CASQ2 leak → DADs RYR2_CPVT
Atrial fibrillation Courtemanche/Nygren atrial cell → 2D/3D tissue; rotor/ablation studies Atrial_Fibrillation, Familial_Atrial_Fibrillation
HCM / DCM Electromechanical FEM + cell remodeling (ToR-ORd validated in HCM) Hypertrophic_Cardiomyopathy, Dilated_Cardiomyopathy
Heart failure Cell remodeling (Grandi/Shannon–Bers) + organ electromechanics Heart_Failure
MI / ischemia Tissue reaction-diffusion with regional hyperkalemia/acidosis, scar geometry Myocardial_Infarction

The CiPA initiative

CiPA (Comprehensive in vitro Proarrhythmia Assay; FDA/HESI/CSRC) replaces the hERG-plus-QT paradigm with mechanistic in-vitro ion-channel data + an in-silico cell model + hiPSC-CM confirmation. The consensus in-silico model is the ORd model with a Markov IKr model capturing dynamic drug–hERG binding ("IKr-dynamic ORd"), optimized in Dutta 2017; the qNet metric stratifies high/intermediate/low TdP risk.

Repositories, databases, and software

Resource What it is Identifiers Landmark
CellML / Physiome Model Repository (PMR2) Curated CellML cell/tissue models; hosts nearly all named cardiac cell models models.cellml.org/e/<id> exposures (ORd = e/71, ToR-ORd = e/5f1) Yu 2011 (PMID:21216774)
BioModels EBI repository (mostly SBML) BIOMD0000000xxx (curated) Malik-Sheriff 2020 (PMID:31701150)
openCARP Open-source cardiac EP simulator (mono/bidomain); successor to CARP opencarp.org Plank 2021 (PMID:34171774)
Chaste C++ library for cardiac EP; strong CellML integration github.com/Chaste Mirams 2013 (PMID:23516352)
FDA/CiPA Reference implementation of the IKr-dynamic ORd + qNet pipeline github.com/FDA/CiPA Dutta 2017 (PMID:28878692)
ToR-ORd Model code (MATLAB + CellML), validation pipeline github.com/jtmff/torord Tomek 2019 (PMID:31868580)
Cardiac Atlas Project Imaging + statistical shape/motion atlases of normal/pathological hearts cardiacatlas.org Fonseca 2011 (PMID:21737439)

Identifier conventions: BioModels BIOMD0000000###; Physiome/CellML human-readable exposure URIs (models.cellml.org/e/<id>). dismech's schema declares a biomodels: prefix; there is no physiome:/cellml: prefix yet, so CellML models are referenced by full repository_url.

Landmark papers with verified PMIDs

Single-cell electrophysiology lineage

Model Citation PMID
Noble Purkinje Noble D. J Physiol. 1962;160(2):317–352. 14480151
Beeler–Reuter Beeler GW, Reuter H. J Physiol. 1977;268(1):177–210. 874889
Luo–Rudy I Luo CH, Rudy Y. Circ Res. 1991;68(6):1501–1526. 1709839
Luo–Rudy II (dynamic) Luo CH, Rudy Y. Circ Res. 1994;74(6):1071–1096. 7514509
Human atrial (Courtemanche) Courtemanche M, Ramirez RJ, Nattel S. Am J Physiol. 1998;275(1):H301–H321. 9688927 (search-confirmed)
Human atrial (Nygren) Nygren A, et al. Circ Res. 1998;82(1):63–81. 9440706 (search-confirmed)
Shannon–Bers Ca²⁺ Shannon TR, et al. Biophys J. 2004;87(5):3351–3371. 15347581
ten Tusscher 2004 ten Tusscher KHWJ, et al. Am J Physiol Heart Circ Physiol. 2004;286(4):H1573–H1589. 14656705
ten Tusscher–Panfilov 2006 Am J Physiol Heart Circ Physiol. 2006;291(3):H1088–H1100. 16565318
ten Tusscher 2006 (efficient) Phys Med Biol. 2006;51(23):6141–6156. 17110776
Grandi–Pasqualini–Bers J Mol Cell Cardiol. 2010;48(1):112–121. 19835882
O'Hara–Rudy (ORd) O'Hara T, Virág L, Varró A, Rudy Y. PLoS Comput Biol. 2011;7(5):e1002061. 21637795
ToR-ORd Tomek J, et al. eLife. 2019;8:e48890. 31868580

hiPSC-CM models (Paci)

Citation PMID
Paci M, et al. Ann Biomed Eng. 2013;41(11):2334–2348. 23722932
Paci M, et al. Biophys J. 2020;118(10):2596–2611. (all-optical) 32298635 (search-confirmed)

CiPA / drug-induced arrhythmia

Citation PMID
Sager PT, et al. Am Heart J. 2014;167(3):292–300. (CiPA rationale) 24576511
Colatsky T, et al. J Pharmacol Toxicol Methods. 2016;81:15–20. (CiPA update) 27282641
Dutta S, et al. Front Physiol. 2017;8:616. (IKr-dynamic ORd optimization) 28878692 (corrigendum = 29230183)

Tissue/organ methods and review

Citation PMID
Clayton RH, et al. Prog Biophys Mol Biol. 2011;104(1–3):22–48. (mono/bidomain review) 20553746

Mitochondrial cardiac bioenergetics (Cortassa / O'Rourke)

Citation PMID
Cortassa S, et al. Biophys J. 2003;84(4):2734–2755. 12668482
Cortassa S, et al. Biophys J. 2006;91(4):1564–1589. (ECME) 16679365
Wei AC, et al. Biophys J. 2009;96(9):3510–3524. 19289071 (search-confirmed)

Curation notes / how to extend

  • Use model_type: KINETIC for ODE cell models; PHYSIOLOGICAL or DIGITAL_TWIN for organ/whole-heart PDE and electromechanical models. There is no dedicated "PDE" enum value.
  • Map state-variable outputs to HP phenotypes via ModelVariableDescriptor thresholds. Treat thresholds as model-calibrated illustrations, not clinical reference values (e.g. APD90 is the cellular correlate of the QT interval, not a direct QT measurement) and say so in the variable/mapping description, exactly as the CKD-MBD entry does.
  • All model evidence uses evidence_source: COMPUTATIONAL with an exact-quote snippet from the model paper's abstract; validate with the standard reference and term validators before committing.
  • Next candidates in priority order: RYR2_CPVT (Shannon–Bers/Grandi Ca²⁺-leak DAD models), Atrial_Fibrillation (Courtemanche), Brugada_Syndrome (reduced-I_Na tissue models), Hypertrophic_Cardiomyopathy (ToR-ORd, validated in HCM), Heart_Failure (Grandi remodeling).