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1
Inheritance
3
Pathophys.
2
Phenotypes
3
Pathograph
1
Medical Actions
👪

Inheritance

1
Compound Inheritance (Null Allele plus Hypomorphic Haplotype)
A rare TBX6 null allele (often a 16p11.2 deletion) in trans with a common hypomorphic TBX6 risk haplotype; a heterozygous null alone is insufficient, so the phenotype reflects compound TBX6 dosage insufficiency.
Show evidence (1 reference)
PMID:25564734 SUPPORT Human Clinical
"Compound inheritance of a rare null mutation and a hypomorphic allele of TBX6 accounted for up to 11% of congenital scoliosis cases in the"
Establishes the compound (null-plus-hypomorphic) TBX6 inheritance model and its substantial contribution to congenital scoliosis.

Pathophysiology

3
TBX6 Dosage Insufficiency at the Determination Wavefront
TBX6 is a T-box transcription factor of the paraxial mesoderm and determination wavefront required for somite formation. Compound inheritance of a rare null allele and a common hypomorphic haplotype lowers functional TBX6 below the threshold needed to drive normal segmentation; a heterozygous null allele alone is insufficient, indicating a dosage-dependent wavefront defect.
Presomitic (Paraxial) Mesoderm Cell CL:0011007
TBX6 hgnc:11605
Somitogenesis GO:0001756 ⚠ ABNORMAL
Show evidence (2 references)
PMID:25564734 SUPPORT Human Clinical
"heterozygous TBX6 null mutation is insufficient to cause congenital scoliosis"
Shows a single null allele is insufficient, establishing the dosage-dependent TBX6 wavefront mechanism.
PMID:25564734 SUPPORT In Vitro
"In vitro functional assays suggested that the risk haplotype is a hypomorphic allele."
Confirms the common risk haplotype is hypomorphic, the second hit lowering TBX6 dosage. Evidence source is IN_VITRO (functional assays).
Disrupted Paraxial Mesoderm Somite Formation
Sub-threshold TBX6 activity corrupts the formation and boundary positioning of somites in the presomitic mesoderm, mis-specifying the metameric template from which the vertebrae derive.
Presomitic (Paraxial) Mesoderm Cell CL:0011007
Segmentation GO:0035282 ⚠ ABNORMAL
Show evidence (1 reference)
PMID:17965051 SUPPORT Model Organism
"The FGF pathway establishes a posterior-to-anterior signaling gradient in the presomitic mesoderm (PSM), which controls cell maturation and is involved in the positioning of segmental boundaries."
Describes the PSM determination wavefront positioning segmental boundaries, the somite-formation step disrupted by reduced TBX6 dosage. Evidence source is MODEL_ORGANISM (mouse).
Hemivertebrae and Congenital Scoliosis
The somite-formation defect manifests as vertebral malsegmentation — characteristically hemivertebrae — that produces a congenital lateral curvature of the spine. Because the vertebrae are serially homologous somite derivatives, one or more segments are malformed.
Chondrocyte CL:0000138 Osteoblast CL:0000062
Embryonic Skeletal System Morphogenesis GO:0048704 ⚠ ABNORMAL
Show evidence (2 references)
PMID:25564734 SUPPORT Human Clinical
"Hemivertebrae are characteristic of TBX6-associated congenital scoliosis."
Identifies hemivertebrae as the characteristic vertebral malsegmentation of TACS, the module consequence.
PMID:25564734 SUPPORT Human Clinical
"Congenital scoliosis is a common type of vertebral malformation."
Frames congenital scoliosis as a vertebral malformation, the clinical endpoint of the malsegmentation.

Pathograph

Use the checkboxes to hide or show graph categories. Hover nodes for evidence and cross-linked metadata.
Pathograph: causal mechanism network for TBX6-Associated Congenital Scoliosis Interactive directed graph showing how pathophysiology mechanisms, phenotypes, genetic factors and variants, experimental models, environmental triggers, and treatments relate through causal and linked edges.

Phenotypes

2
Hemivertebrae HP:0002937
Show evidence (1 reference)
PMID:25564734 SUPPORT Human Clinical
"Hemivertebrae are characteristic of TBX6-associated congenital scoliosis."
Hemivertebrae are the characteristic vertebral defect of TACS.
Congenital Scoliosis HP:0002650
Show evidence (1 reference)
PMID:25564734 SUPPORT Human Clinical
"Congenital scoliosis is a common type of vertebral malformation."
TACS presents as congenital scoliosis, a vertebral malformation.
💊

Medical Actions

1
Scoliosis Surgical Management
Action: orthopedic surgical procedure Ontology label: Orthopedic Surgical Procedure NCIT:C16186
Management of congenital scoliosis ranges from observation and bracing to growth-friendly instrumentation or corrective/fusion surgery for progressive curves; hemivertebra resection may be indicated for focal deformity.
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Source YAML

click to show
name: TBX6-Associated Congenital Scoliosis
creation_date: "2026-06-29T00:00:00Z"
category: Mendelian
description: >-
  TBX6-associated congenital scoliosis (TACS) is a congenital axial-skeleton
  malformation in which vertebral mis-segmentation — characteristically
  hemivertebrae — produces congenital scoliosis. It is the most common defined
  genetic cause of congenital scoliosis and follows a distinctive compound
  inheritance model: a rare loss-of-function (null) TBX6 allele — frequently a
  recurrent 16p11.2 deletion encompassing TBX6 — in trans with a common
  hypomorphic TBX6 risk haplotype, so that TBX6 dosage falls below the threshold
  required for normal somite formation while a heterozygous null alone is
  insufficient. TBX6 is a T-box transcription factor of the paraxial mesoderm /
  determination wavefront that is required to form somites, the serially
  repeated precursors of the vertebrae. TACS is filed in OMIM/MONDO under
  spondylocostal dysostosis 5 (OMIM:122600) but is clinically a milder, often
  more localized congenital scoliosis than classic recessive spondylocostal
  dysostosis. It conforms to the axial-skeleton serial-homology module across
  its wavefront, somite-formation, and vertebral-malsegmentation nodes.
disease_term:
  preferred_term: TBX6-associated congenital scoliosis
  term:
    id: MONDO:0007389
    label: spondylocostal dysostosis 5
parents:
- Vertebral Malformation Disorders
inheritance:
- name: Compound Inheritance (Null Allele plus Hypomorphic Haplotype)
  description: >-
    A rare TBX6 null allele (often a 16p11.2 deletion) in trans with a common
    hypomorphic TBX6 risk haplotype; a heterozygous null alone is insufficient,
    so the phenotype reflects compound TBX6 dosage insufficiency.
  evidence:
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: Compound inheritance of a rare null mutation and a hypomorphic allele of TBX6 accounted for up to 11% of congenital scoliosis cases in the
    explanation: >-
      Establishes the compound (null-plus-hypomorphic) TBX6 inheritance model
      and its substantial contribution to congenital scoliosis.
prevalence:
- population: Sporadic congenital scoliosis (Han Chinese series)
  percentage: ~11%
  notes: >-
    TBX6 null variants plus the common hypomorphic haplotype accounted for up to
    11% of sporadic congenital scoliosis cases in the original Han Chinese
    series; TACS is the most common defined monogenic-dosage cause of congenital
    scoliosis identified to date.
  evidence:
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: we identified a total of 17 heterozygous TBX6 null mutations in the 161
    explanation: >-
      Reports 17 TBX6 null mutations among 161 persons with sporadic congenital
      scoliosis (~11%).
pathophysiology:
- name: TBX6 Dosage Insufficiency at the Determination Wavefront
  conforms_to: "axial_segmentation_serial_homology#Segmentation Clock and Wavefront Dysfunction"
  description: >-
    TBX6 is a T-box transcription factor of the paraxial mesoderm and
    determination wavefront required for somite formation. Compound inheritance
    of a rare null allele and a common hypomorphic haplotype lowers functional
    TBX6 below the threshold needed to drive normal segmentation; a heterozygous
    null allele alone is insufficient, indicating a dosage-dependent wavefront
    defect.
  role: trigger
  genes:
  - preferred_term: TBX6
    term:
      id: hgnc:11605
      label: TBX6
  cell_types:
  - preferred_term: Presomitic (Paraxial) Mesoderm Cell
    term:
      id: CL:0011007
      label: paraxial cell
  biological_processes:
  - preferred_term: Somitogenesis
    term:
      id: GO:0001756
      label: somitogenesis
    modifier: ABNORMAL
  evidence:
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: heterozygous TBX6 null mutation is insufficient to cause congenital scoliosis
    explanation: >-
      Shows a single null allele is insufficient, establishing the
      dosage-dependent TBX6 wavefront mechanism.
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: In vitro functional assays suggested that the risk haplotype is a hypomorphic allele.
    explanation: >-
      Confirms the common risk haplotype is hypomorphic, the second hit lowering
      TBX6 dosage. Evidence source is IN_VITRO (functional assays).
  downstream:
  - target: Disrupted Paraxial Mesoderm Somite Formation
    description: Reduced TBX6 dosage impairs somite formation in the paraxial mesoderm.
    causal_link_type: INDIRECT_KNOWN_INTERMEDIATES
    intermediate_mechanisms:
    - sub-threshold TBX6 transcriptional output in the presomitic mesoderm
- name: Disrupted Paraxial Mesoderm Somite Formation
  conforms_to: "axial_segmentation_serial_homology#Disrupted Somite Boundary Formation"
  description: >-
    Sub-threshold TBX6 activity corrupts the formation and boundary positioning
    of somites in the presomitic mesoderm, mis-specifying the metameric template
    from which the vertebrae derive.
  role: central_effector
  cell_types:
  - preferred_term: Presomitic (Paraxial) Mesoderm Cell
    term:
      id: CL:0011007
      label: paraxial cell
  biological_processes:
  - preferred_term: Segmentation
    term:
      id: GO:0035282
      label: segmentation
    modifier: ABNORMAL
  evidence:
  - reference: PMID:17965051
    supports: SUPPORT
    evidence_source: MODEL_ORGANISM
    snippet: The FGF pathway establishes a posterior-to-anterior signaling gradient in the presomitic mesoderm (PSM), which controls cell maturation and is involved in the positioning of segmental boundaries.
    explanation: >-
      Describes the PSM determination wavefront positioning segmental
      boundaries, the somite-formation step disrupted by reduced TBX6 dosage.
      Evidence source is MODEL_ORGANISM (mouse).
  downstream:
  - target: Hemivertebrae and Congenital Scoliosis
    description: Mis-formed somites yield hemivertebrae and a curved spine.
    causal_link_type: INDIRECT_KNOWN_INTERMEDIATES
    intermediate_mechanisms:
    - asymmetric/incomplete vertebral body formation from mis-specified somites
- name: Hemivertebrae and Congenital Scoliosis
  conforms_to: "axial_segmentation_serial_homology#Vertebral and Costal Malsegmentation"
  description: >-
    The somite-formation defect manifests as vertebral malsegmentation —
    characteristically hemivertebrae — that produces a congenital lateral
    curvature of the spine. Because the vertebrae are serially homologous somite
    derivatives, one or more segments are malformed.
  role: consequence
  cell_types:
  - preferred_term: Chondrocyte
    term:
      id: CL:0000138
      label: chondrocyte
  - preferred_term: Osteoblast
    term:
      id: CL:0000062
      label: osteoblast
  biological_processes:
  - preferred_term: Embryonic Skeletal System Morphogenesis
    term:
      id: GO:0048704
      label: embryonic skeletal system morphogenesis
    modifier: ABNORMAL
  evidence:
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: Hemivertebrae are characteristic of TBX6-associated congenital scoliosis.
    explanation: >-
      Identifies hemivertebrae as the characteristic vertebral malsegmentation
      of TACS, the module consequence.
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: Congenital scoliosis is a common type of vertebral malformation.
    explanation: >-
      Frames congenital scoliosis as a vertebral malformation, the clinical
      endpoint of the malsegmentation.
phenotypes:
- name: Hemivertebrae
  description: >-
    Hemivertebrae (incomplete, wedge-shaped vertebral bodies) are the
    characteristic vertebral malsegmentation of TACS.
  phenotype_term:
    preferred_term: Hemivertebrae
    term:
      id: HP:0002937
      label: Hemivertebrae
  evidence:
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: Hemivertebrae are characteristic of TBX6-associated congenital scoliosis.
    explanation: Hemivertebrae are the characteristic vertebral defect of TACS.
- name: Congenital Scoliosis
  description: >-
    Congenital lateral spinal curvature resulting from the vertebral
    malsegmentation.
  phenotype_term:
    preferred_term: Scoliosis
    term:
      id: HP:0002650
      label: Scoliosis
  evidence:
  - reference: PMID:25564734
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: Congenital scoliosis is a common type of vertebral malformation.
    explanation: TACS presents as congenital scoliosis, a vertebral malformation.
treatments:
- name: Scoliosis Surgical Management
  description: >-
    Management of congenital scoliosis ranges from observation and bracing to
    growth-friendly instrumentation or corrective/fusion surgery for progressive
    curves; hemivertebra resection may be indicated for focal deformity.
  treatment_term:
    preferred_term: orthopedic surgical procedure
    term:
      id: NCIT:C16186
      label: Orthopedic Surgical Procedure