Microvillus Inclusion Disease

Microvillus inclusion disease (MVID) is an ultra-rare autosomal recessive congenital enteropathy and the prototype of the "defective enterocyte trafficking and polarity" class of congenital diarrheas and enteropathies (CODEs). Biallelic loss-of-function variants in MYO5B, which encodes the unconventional actin-based motor myosin Vb, disable apical recycling-endosome traffic in the polarized enterocyte. Apical transporters and brush-border components are no longer delivered to or retained at the apical membrane; instead the brush border is lost and internalized microvilli accumulate inside the cell as the pathognomonic intracytoplasmic microvillus inclusions, with subapical accumulation of periodic-acid-Schiff-positive secretory granules. Affected neonates have profuse, life-threatening watery diarrhea that begins in the first days of life (early-onset form) or within the first months (late-onset form) and, critically, does NOT remit on fasting or on removal of any dietary constituent - the feature that separates MVID from the substrate-dependent diet-induced congenital diarrheas. Children are dependent on parenteral nutrition from birth, and the natural history is dominated by the complications of that dependence: intestinal-failure-associated liver disease, catheter sepsis, growth failure and renal impairment. Small-bowel transplantation remains the only definitive treatment. Because myosin Vb also organizes apical traffic at the hepatocyte canalicular membrane, MYO5B variants additionally cause a low-GGT intrahepatic cholestasis (PFIC10) that may occur with or without the enteropathy, making MVID a two-organ disease of one shared trafficking mechanism rather than a purely intestinal disorder.

Ask OpenScientist

Ask a research question about Microvillus Inclusion Disease. OpenScientist will conduct autonomous deep research using the Disorder Mechanisms Knowledge Base and PubMed literature (typically 10-30 minutes).

Submitting...

Do not include personal health information in your question. Questions and results are cached in your browser's local storage.

1
Inheritance
5
Pathophys.
5
Phenotypes
7
Pathograph
2
Genes
2
Medical Actions
👪

Inheritance

1
Autosomal recessive inheritance HP:0000007
MVID is inherited in an autosomal recessive manner. Reported cohorts are strongly enriched for consanguineous families, particularly of Mediterranean and Middle Eastern origin, and affected individuals are typically homozygous for a founder or private MYO5B variant.
Autosomal recessive inheritance
Show evidence (1 reference)
PMID:21407114 SUPPORT Human Clinical
"Almost half of the children were from consanguineous families from the Mediterranean area."
Documents the consanguinity enrichment expected of a recessive disorder in the largest single-centre MVID cohort reported.

Pathophysiology

5
MYO5B Loss and Failure of Apical Recycling-Endosome Traffic
The initiating molecular lesion. Myosin Vb is an actin-based motor that moves apical recycling endosomes along the subapical actin network of the polarized enterocyte and, with its RAB8/RAB11 partners and the apical SNARE syntaxin-3, delivers their cargo to the apical plasma membrane. Biallelic loss of the motor - or of syntaxin-3, the receiving SNARE one step downstream - arrests this delivery step. Cargo destined for the apical surface is retained in the subapical endosomal compartment, and, as the original gene-discovery work showed by the mislocalization of transferrin receptor, basolateral trafficking is disturbed as well, so the defect is one of general epithelial polarity rather than of a single apical protein.
Small-intestinal enterocyte CL:0000584 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Small-intestinal enterocyte, annotated with enterocyte (CL:0000584). CL:0000584 is a cell type from the Cell Ontology.
Apical delivery of vesicular cargo GO:0016192 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves Apical delivery of vesicular cargo, annotated with vesicle-mediated transport (GO:0016192), qualified as loss of function. GO:0016192 is a biological process from the Gene Ontology. ⇓ LOSS OF FUNCTION Establishment of enterocyte apical-basal polarity GO:0045198 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Establishment of enterocyte apical-basal polarity, annotated with establishment of epithelial cell apical/basal polarity (GO:0045198). GO:0045198 is a biological process from the Gene Ontology. ⚠ ABNORMAL
Apical recycling endosome GO:0055037 Gene Ontology (GO) Relation: this pathophysiological event involves this cellular component This pathophysiological event involves Apical recycling endosome, annotated with recycling endosome (GO:0055037). GO:0055037 is a cellular component from the Gene Ontology.
Small intestine UBERON:0002108 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Small intestine (UBERON:0002108). UBERON:0002108 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (1 reference)
PMID:18724368 SUPPORT Human Clinical
"mislocalization of transferrin receptor in MVID enterocytes suggests that MYO5B deficiency causes defective trafficking of apical and basolateral proteins in MVID"
Establishes the trigger as a general defect of polarized membrane traffic - affecting basolateral cargo (transferrin receptor) as well as apical cargo - rather than loss of one apical transporter.
Brush-Border Loss and Intracellular Microvillus Inclusion Formation
The defining structural lesion and the diagnostic hallmark. Without myosin-Vb-dependent apical delivery, the enterocyte cannot build or maintain its brush border: apical microvilli are sparse or absent over the villus surface. Microvilli that do form are internalized into vacuolar structures within the cytoplasm, producing the pathognomonic microvillus inclusions - intracellular vacuoles lined by fully formed microvilli - accompanied by accumulation of periodic-acid-Schiff-positive granules in the subapical cytoplasm. Villous atrophy is present, so unlike the diet-induced congenital diarrheas the villus-to-crypt ratio is abnormal, which is what places MVID in the enterocyte-structural rather than the digestive-absorptive CODE class.
Small-intestinal enterocyte CL:0000584 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Small-intestinal enterocyte, annotated with enterocyte (CL:0000584). CL:0000584 is a cell type from the Cell Ontology.
Microvillus assembly at the apical surface GO:0030033 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves decreased Microvillus assembly at the apical surface, annotated with microvillus assembly (GO:0030033). GO:0030033 is a biological process from the Gene Ontology. ↓ DECREASED Apical membrane protein delivery GO:0072659 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves decreased Apical membrane protein delivery, annotated with protein localization to plasma membrane (GO:0072659). GO:0072659 is a biological process from the Gene Ontology. ↓ DECREASED
Microvillus GO:0005902 Gene Ontology (GO) Relation: this pathophysiological event involves this cellular component This pathophysiological event involves Microvillus (GO:0005902). GO:0005902 is a cellular component from the Gene Ontology.
Show evidence (1 reference)
PMID:18724368 SUPPORT Human Clinical
"MVID is characterized by lack of microvilli on the surface of enterocytes and occurrence of intracellular vacuolar structures containing microvilli."
States both halves of this node: absence of surface microvilli and the intracellular microvillus-containing vacuoles that define the disease histologically.
Loss of Apical Absorptive and Ion-Transport Capacity
The functional consequence of losing the apical membrane. The brush border is the entire absorptive interface of the enterocyte, carrying the disaccharidases, the nutrient carriers and the sodium-coupled ion exchangers that drive net fluid absorption. Its loss, together with failure to deliver and retain apical transporters, abolishes absorptive capacity across the board rather than for one substrate. The resulting diarrhea is therefore non-selective and, distinctively, persists during complete bowel rest - fluid loss continues when nothing is being fed, because the defect is in the absorptive surface itself and not in the handling of any ingested nutrient.
Small-intestinal enterocyte CL:0000584 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Small-intestinal enterocyte, annotated with enterocyte (CL:0000584). CL:0000584 is a cell type from the Cell Ontology.
Intestinal absorption GO:0050892 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves decreased Intestinal absorption (GO:0050892). GO:0050892 is a biological process from the Gene Ontology. ↓ DECREASED
Show evidence (2 references)
PMID:29654747 SUPPORT Other
"diminution of brush-border microvillar markers, including ezrin, villin, phospho-ERM, NHE3, and DPP-IV"
Names the specific apical machinery lost at this node, including the sodium/hydrogen exchanger NHE3 and the brush-border peptidase DPP-IV - direct evidence that losing the surface removes transport and digestive capacity together rather than one selective step. Evidence source is OTHER because this is a review.
PMID:33374714 SUPPORT Other
"Along with microvillus inclusion disease, a severe disorder belonging to the defective enterocyte trafficking and polarity category of CODEs is congenital tufting enteropathy (CTE)."
Supporting context only: places MVID in the defective enterocyte trafficking and polarity CODE class. It establishes the classification, not the absorptive-capacity claim, which the preceding citation carries.
Intractable Secretory Diarrhea and Lifelong Intestinal Failure
The clinical output. Profuse watery diarrhea of very high volume begins in the first days of life in the early-onset form, causing severe dehydration, hyponatremia and metabolic acidosis that require immediate intravenous replacement. Because enteral feeding cannot sustain the child, total parenteral nutrition is required from the neonatal period and, unlike in most causes of intestinal failure, enteral autonomy is essentially never achieved. The disease course is then dominated by the complications of lifelong parenteral nutrition - intestinal-failure-associated liver disease, catheter-related sepsis, growth failure, developmental delay, renal impairment and metabolic bone disease.
Small intestine UBERON:0002108 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Small intestine (UBERON:0002108). UBERON:0002108 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (2 references)
PMID:21407114 SUPPORT Human Clinical
"All of the patients completely depended on PN."
Documents the absolute and universal parenteral-nutrition dependence that defines the clinical output of this node in a 24-patient cohort.
PMID:21407114 SUPPORT Human Clinical
"Before or without SBTx, growth failure was common (mean height -2.5 standard deviations [SD]), as was developmental delay (12/24), liver (20/22 with fibrosis) or kidney disease (3/23 with moderate renal insufficiency), and osteoporosis (6/24)."
Quantifies the parenteral-nutrition-dependent complication burden - growth failure, developmental delay, liver fibrosis, renal impairment and osteoporosis - that dominates the natural history.
Canalicular Trafficking Failure and Low-GGT Intrahepatic Cholestasis
The hepatic arm of the same trafficking lesion. Myosin Vb organizes apical membrane traffic in the hepatocyte just as it does in the enterocyte, and the hepatocyte's apical membrane is the bile canaliculus. Loss of the motor mislocalizes the canalicular bile-salt export pump and disorganizes the canaliculus, producing a low-gamma-glutamyl-transferase intrahepatic cholestasis (PFIC10). This arm can occur together with the enteropathy or, with certain MYO5B genotypes, in isolation without any intestinal disease, which is why MYO5B disease is best understood as one trafficking defect expressed in two polarized epithelia. It is mechanistically distinct from - and must not be conflated with - the intestinal-failure-associated liver disease that parenteral nutrition itself causes in these same children.
Liver UBERON:0002107 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Liver (UBERON:0002107). UBERON:0002107 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (3 references)
PMID:18724368 SUPPORT Human Clinical
"MYO5B deficiency causes defective trafficking of apical and basolateral proteins"
Supports the shared trafficking mechanism underlying this arm, which in the hepatocyte applies to the canalicular (apical) membrane. This citation establishes the trafficking defect, not the hepatic phenotype itself, which is evidenced separately below.
PMID:27532546 SUPPORT Human Clinical
"Some patients with microvillus inclusion disease due to myosin 5B (MYO5B) mutations may develop cholestasis characterized by a progressive familial intrahepatic cholestasis-like phenotype with normal serum gamma-glutamyl transferase activity."
Establishes the hepatic phenotype of this node in patients who also have MVID - a low-GGT, PFIC-like intrahepatic cholestasis attributable to MYO5B deficiency.
PMID:27532546 SUPPORT Human Clinical
"These data show that MYO5B deficiency may lead to isolated cholestasis and that MYO5B should be considered as an additional progressive familial intrahepatic cholestasis gene."
Supports the dissociation claim central to this node: MYO5B deficiency can produce cholestasis in the absence of any intestinal disease, which is why the hepatic arm is modelled as a parallel effector of the same trafficking lesion rather than as a complication of the enteropathy.

Pathograph

Use the checkboxes to hide or show graph categories. Hover nodes for evidence and cross-linked metadata.
Pathograph: causal mechanism network for Microvillus Inclusion Disease Interactive directed graph showing how pathophysiology mechanisms, phenotypes, genetic factors and variants, experimental models, environmental triggers, and treatments relate through causal and linked edges.

Phenotypes

5
Digestive 2
Intractable Watery Diarrhea OBLIGATE HP:0002014 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Intractable watery diarrhea, annotated with Diarrhea (HP:0002014), qualified as temporality chronic. HP:0002014 is a phenotype from the Human Phenotype Ontology.
Temporal: CHRONIC
Show evidence (1 reference)
PMID:21407114 SUPPORT Human Clinical
"Microvillous inclusion disease (MVID) is a rare congenital enterocyte disorder causing severe diarrhea and intestinal failure."
States severe diarrhea as the defining clinical feature of the disorder.
Cholestasis HP:0001396 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Cholestasis (HP:0001396). HP:0001396 is a phenotype from the Human Phenotype Ontology.
Curated from clinical description of MYO5B-related disease. Liver fibrosis in the cited MVID cohort (20/22) is attributable to parenteral nutrition rather than to the MYO5B canalicular defect, so that cohort figure is deliberately not used as evidence for this phenotype and no frequency band is asserted.
Metabolism 1
Dehydration HP:0001944 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Dehydration (HP:0001944), qualified as temporality acute. HP:0001944 is a phenotype from the Human Phenotype Ontology.
Temporal: ACUTE
Show evidence (1 reference)
PMID:29654747 SUPPORT Other
"Patients with MVID present with severe watery, electrolyte transport-related diarrhea and dehydration with failure to thrive beginning in the first days or weeks of life."
States dehydration as a presenting feature of MVID specifically, with its neonatal onset, rather than as generic guidance about inferring diarrhea. Evidence source is OTHER because this is a review.
Growth 1
Failure to Thrive VERY_FREQUENT HP:0001508 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Failure to thrive (HP:0001508), qualified as course progressive. HP:0001508 is a phenotype from the Human Phenotype Ontology.
Course: PROGRESSIVE
Show evidence (1 reference)
PMID:21407114 SUPPORT Human Clinical
"Before or without SBTx, growth failure was common (mean height -2.5 standard deviations [SD])"
Directly quantifies growth failure in the cohort, supporting both the phenotype and its VERY_FREQUENT frequency band.
Other 1
Villous Atrophy HP:0011473 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Villous atrophy (HP:0011473). HP:0011473 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:29654747 SUPPORT Other
"Other pathological hallmarks of the disease include villus atrophy, abnormal subapical periodic acid-Schiff/CD10 staining in enterocytes, inclusion staining with antibodies against ezrin or villin"
States villus atrophy directly, together with the subapical periodic-acid-Schiff/CD10 staining and inclusion staining that accompany it - the mucosal architecture claim this phenotype makes, as distinct from the microvillar ultrastructure cited on the pathophysiology node.
🧬

Genetic Associations

2
MYO5B loss-of-function variants (Causative)
Gene: MYO5B hgnc:7603 HUGO Gene Nomenclature Committee (hgnc) Relation: this disease-associated gene is this gene This disease-associated gene is MYO5B (hgnc:7603). hgnc:7603 is a gene from the HUGO Gene Nomenclature Committee. relationship_type: CAUSATIVE variant_origin: GERMLINE
Show evidence (1 reference)
PMID:18724368 SUPPORT Human Clinical
"Following homozygosity mapping in a single kindred, we identified nonsense and missense mutations in MYO5B, encoding type Vb myosin motor protein, in individuals with microvillus inclusion disease (MVID)."
The original gene-discovery report establishing biallelic MYO5B variants as the cause of MVID.
STX3 variants (MVID variant form) (Causative)
Gene: STX3 hgnc:11438 HUGO Gene Nomenclature Committee (hgnc) Relation: this disease-associated gene is this gene This disease-associated gene is STX3 (hgnc:11438). hgnc:11438 is a gene from the HUGO Gene Nomenclature Committee. relationship_type: CAUSATIVE variant_origin: GERMLINE
Show evidence (1 reference)
PMID:35769957 SUPPORT Human Clinical
"Newer reports of mutations resulting in truncated syntaxin 3 (STX3) and Munc18-2 (STXBP2) proteins have been elucidated as causative."
Establishes STX3 (and STXBP2/Munc18-2) as causative loci for MVID alongside MYO5B, supporting the variant-form genetic block. The same sentence names STXBP2, the third locus in the apical-delivery pathway.
💊

Medical Actions

2
Total Parenteral Nutrition
Action: Total Parenteral NutritionNCI Thesaurus (NCIT) Relation: this treatment is this clinical intervention This treatment is Total Parenteral Nutrition (NCIT:C29484). NCIT:C29484 is a clinical intervention from the NCI Thesaurus. NCIT:C29484
Lifelong total parenteral nutrition is required from the neonatal period, because the absorptive surface is absent and enteral autonomy is essentially never achieved. It is supportive, not disease-modifying: it sustains the child while the underlying trafficking defect persists, and it carries its own substantial morbidity (intestinal-failure-associated liver disease, catheter sepsis, metabolic bone disease, renal impairment). Management in an expert intestinal-failure centre measurably improves outcome.
Mechanism Target:
BYPASSES Intractable Secretory Diarrhea and Lifelong Intestinal Failure — Parenteral nutrition bypasses the non-functional intestinal absorptive surface entirely, delivering nutrition intravenously; it does not act on the trafficking defect or restore the brush border.
Show evidence (1 reference)
PMID:21407114 SUPPORT Human Clinical
"PN in MVID is difficult to manage and requires expertise."
Supports parenteral nutrition as the mainstay of management and records that outcome depends on the expertise with which it is delivered.
Small-Bowel Transplantation
Action: Small Bowel TransplantationNCI Thesaurus (NCIT) Relation: this treatment is this clinical intervention This treatment is Small Bowel Transplantation (NCIT:C157985). NCIT:C157985 is a clinical intervention from the NCI Thesaurus. NCIT:C157985
Small-bowel transplantation, isolated or combined with liver transplantation when intestinal-failure-associated liver disease has advanced, is the only definitive treatment. It replaces the defective epithelium with donor intestine carrying two functional MYO5B alleles. In a 24-patient cohort, survival was 77% with transplantation versus 63% without, and catch-up growth followed transplantation in some children.
Mechanism Target:
RESTORES Loss of Apical Absorptive and Ion-Transport Capacity — Replacing the intestine with genotypically normal donor epithelium restores an intact brush border and absorptive capacity, the only intervention that addresses the effector node rather than compensating for it.
Show evidence (2 references)
PMID:21407114 SUPPORT Human Clinical
"SBTx, despite being complicated, remains the only hope to improve the quality of life and long-term prognosis of these children."
States that small-bowel transplantation is the only intervention that improves long-term prognosis, supporting it as the definitive treatment.
PMID:21407114 SUPPORT Human Clinical
"Patient survival rates were 63% without SBTx and 77% with SBTx."
Quantifies the survival difference associated with transplantation in the cohort.
🔬

Diagnosis

2
Small-intestinal biopsy with electron microscopy
Duodenal or jejunal biopsy is the diagnostic cornerstone. Light microscopy shows villous atrophy without significant inflammatory infiltrate; periodic acid-Schiff and CD10 staining reveal abnormal subapical granular deposits; electron microscopy demonstrates the pathognomonic intracytoplasmic microvillus inclusions and the sparse or absent apical brush border.
Show evidence (1 reference)
PMID:18724368 SUPPORT Human Clinical
"MVID is characterized by lack of microvilli on the surface of enterocytes and occurrence of intracellular vacuolar structures containing microvilli."
Describes the two ultrastructural findings that establish the diagnosis on electron microscopy.
MYO5B and STX3 sequencing
Molecular confirmation by sequencing of MYO5B and, where MYO5B is negative, STX3. Whole-exome or whole-genome sequencing early in the evaluation of neonatal intractable diarrhea shortens the diagnostic odyssey and can establish the diagnosis before mucosal findings are available.
Show evidence (1 reference)
PMID:29654747 SUPPORT Other
"Whole-exome and genome sequencing in the early stages of evaluation can reduce the time required for a definitive diagnosis of CODEs"
Supports early genomic testing as the recommended diagnostic strategy for congenital diarrheas including MVID. Evidence source is OTHER because this is a review.
{ }

Source YAML

click to show
name: Microvillus Inclusion Disease
creation_date: "2026-08-22T00:00:00Z"
category: Mendelian
synonyms:
- MVID
- microvillous inclusion disease
- congenital microvillus atrophy
- Davidson disease
- diarrhea 2, with microvillus atrophy
description: >-
  Microvillus inclusion disease (MVID) is an ultra-rare autosomal recessive
  congenital enteropathy and the prototype of the "defective enterocyte
  trafficking and polarity" class of congenital diarrheas and enteropathies
  (CODEs). Biallelic loss-of-function variants in MYO5B, which encodes the
  unconventional actin-based motor myosin Vb, disable apical recycling-endosome
  traffic in the polarized enterocyte. Apical transporters and brush-border
  components are no longer delivered to or retained at the apical membrane;
  instead the brush border is lost and internalized microvilli accumulate inside
  the cell as the pathognomonic intracytoplasmic microvillus inclusions, with
  subapical accumulation of periodic-acid-Schiff-positive secretory granules.
  Affected neonates have profuse, life-threatening watery diarrhea that begins in
  the first days of life (early-onset form) or within the first months
  (late-onset form) and, critically, does NOT remit on fasting or on removal of
  any dietary constituent - the feature that separates MVID from the
  substrate-dependent diet-induced congenital diarrheas. Children are dependent
  on parenteral nutrition from birth, and the natural history is dominated by
  the complications of that dependence: intestinal-failure-associated liver
  disease, catheter sepsis, growth failure and renal impairment. Small-bowel
  transplantation remains the only definitive treatment. Because myosin Vb also
  organizes apical traffic at the hepatocyte canalicular membrane, MYO5B
  variants additionally cause a low-GGT intrahepatic cholestasis (PFIC10) that
  may occur with or without the enteropathy, making MVID a two-organ disease of
  one shared trafficking mechanism rather than a purely intestinal disorder.
disease_term:
  preferred_term: microvillus inclusion disease
  term:
    id: MONDO:0009635
    label: microvillus inclusion disease
inheritance:
- name: Autosomal recessive inheritance
  description: >-
    MVID is inherited in an autosomal recessive manner. Reported cohorts are
    strongly enriched for consanguineous families, particularly of Mediterranean
    and Middle Eastern origin, and affected individuals are typically homozygous
    for a founder or private MYO5B variant.
  inheritance_term:
    preferred_term: Autosomal recessive inheritance
    term:
      id: HP:0000007
      label: Autosomal recessive inheritance
  evidence:
  - reference: PMID:21407114
    reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Almost half of the children were from consanguineous families from the
      Mediterranean area.
    explanation: >-
      Documents the consanguinity enrichment expected of a recessive disorder in
      the largest single-centre MVID cohort reported.
genetic:
- name: MYO5B loss-of-function variants
  gene_term:
    preferred_term: MYO5B
    term:
      id: hgnc:7603
      label: MYO5B
  association: Causative
  relationship_type: CAUSATIVE
  variant_origin: GERMLINE
  features: >-
    Biallelic nonsense, frameshift, splice and missense variants in MYO5B,
    encoding the type Vb myosin motor, are the principal cause of MVID. The
    motor couples apical recycling endosomes to the actin cytoskeleton, so its
    loss disrupts trafficking of both apical and basolateral membrane proteins
    in the enterocyte. Genotype partly predicts organ involvement: some MYO5B
    variants produce isolated low-GGT cholestasis without enteropathy, indicating
    that intestinal and hepatic manifestations can dissociate.
  evidence:
  - reference: PMID:18724368
    reference_title: "MYO5B mutations cause microvillus inclusion disease and disrupt epithelial cell polarity."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Following homozygosity mapping in a single kindred, we identified nonsense
      and missense mutations in MYO5B, encoding type Vb myosin motor protein, in
      individuals with microvillus inclusion disease (MVID).
    explanation: >-
      The original gene-discovery report establishing biallelic MYO5B variants as
      the cause of MVID.
- name: STX3 variants (MVID variant form)
  gene_term:
    preferred_term: STX3
    term:
      id: hgnc:11438
      label: STX3
  association: Causative
  relationship_type: CAUSATIVE
  variant_origin: GERMLINE
  features: >-
    Biallelic variants in STX3, encoding the apical SNARE syntaxin-3 that
    receives myosin-Vb-delivered vesicles at the enterocyte apical membrane,
    cause a variant form of MVID that is histologically indistinguishable from
    the MYO5B form. STX3 acts immediately downstream of MYO5B in the same apical
    delivery step, so its loss phenocopies the motor defect - genetic evidence
    that the pathogenic lesion is the apical trafficking pathway itself rather
    than the motor protein specifically.
  evidence:
  - reference: PMID:35769957
    reference_title: "Microvillus Inclusion Disease: A Rare Mutation of STX3 in Exon 9 Causing Fatal Congenital Diarrheal Disease."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Newer reports of mutations resulting in truncated syntaxin 3 (STX3) and
      Munc18-2 (STXBP2) proteins have been elucidated as causative.
    explanation: >-
      Establishes STX3 (and STXBP2/Munc18-2) as causative loci for MVID
      alongside MYO5B, supporting the variant-form genetic block. The same
      sentence names STXBP2, the third locus in the apical-delivery pathway.
pathophysiology:
- name: MYO5B Loss and Failure of Apical Recycling-Endosome Traffic
  conforms_to: "enterocyte_polarity_trafficking_failure#Loss of Apical Delivery Machinery"
  description: >-
    The initiating molecular lesion. Myosin Vb is an actin-based motor that moves
    apical recycling endosomes along the subapical actin network of the polarized
    enterocyte and, with its RAB8/RAB11 partners and the apical SNARE
    syntaxin-3, delivers their cargo to the apical plasma membrane. Biallelic
    loss of the motor - or of syntaxin-3, the receiving SNARE one step
    downstream - arrests this delivery step. Cargo destined for the apical
    surface is retained in the subapical endosomal compartment, and, as the
    original gene-discovery work showed by the mislocalization of transferrin
    receptor, basolateral trafficking is disturbed as well, so the defect is one
    of general epithelial polarity rather than of a single apical protein.
  role: trigger
  biological_scale: MOLECULAR
  cell_types:
  - preferred_term: Small-intestinal enterocyte
    term:
      id: CL:0000584
      label: enterocyte
  biological_processes:
  - preferred_term: Apical delivery of vesicular cargo
    term:
      id: GO:0016192
      label: vesicle-mediated transport
    modifier: LOSS_OF_FUNCTION
  - preferred_term: Establishment of enterocyte apical-basal polarity
    term:
      id: GO:0045198
      label: establishment of epithelial cell apical/basal polarity
    modifier: ABNORMAL
  cellular_components:
  - preferred_term: Apical recycling endosome
    term:
      id: GO:0055037
      label: recycling endosome
  locations:
  - preferred_term: Small intestine
    term:
      id: UBERON:0002108
      label: small intestine
  evidence:
  - reference: PMID:18724368
    reference_title: "MYO5B mutations cause microvillus inclusion disease and disrupt epithelial cell polarity."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      mislocalization of transferrin receptor in MVID enterocytes suggests that
      MYO5B deficiency causes defective trafficking of apical and basolateral
      proteins in MVID
    explanation: >-
      Establishes the trigger as a general defect of polarized membrane traffic -
      affecting basolateral cargo (transferrin receptor) as well as apical cargo
      - rather than loss of one apical transporter.
  downstream:
  - target: Brush-Border Loss and Intracellular Microvillus Inclusion Formation
    causal_link_type: DIRECT
  - target: Canalicular Trafficking Failure and Low-GGT Intrahepatic Cholestasis
    causal_link_type: DIRECT
- name: Brush-Border Loss and Intracellular Microvillus Inclusion Formation
  conforms_to: "enterocyte_polarity_trafficking_failure#Loss of a Functionally Polarized Absorptive Epithelial Surface"
  description: >-
    The defining structural lesion and the diagnostic hallmark. Without
    myosin-Vb-dependent apical delivery, the enterocyte cannot build or maintain
    its brush border: apical microvilli are sparse or absent over the villus
    surface. Microvilli that do form are internalized into vacuolar structures
    within the cytoplasm, producing the pathognomonic microvillus inclusions -
    intracellular vacuoles lined by fully formed microvilli - accompanied by
    accumulation of periodic-acid-Schiff-positive granules in the subapical
    cytoplasm. Villous atrophy is present, so unlike the diet-induced congenital
    diarrheas the villus-to-crypt ratio is abnormal, which is what places MVID in
    the enterocyte-structural rather than the digestive-absorptive CODE class.
  role: central_effector
  biological_scale: CELLULAR
  cell_types:
  - preferred_term: Small-intestinal enterocyte
    term:
      id: CL:0000584
      label: enterocyte
  biological_processes:
  - preferred_term: Microvillus assembly at the apical surface
    term:
      id: GO:0030033
      label: microvillus assembly
    modifier: DECREASED
  - preferred_term: Apical membrane protein delivery
    term:
      id: GO:0072659
      label: protein localization to plasma membrane
    modifier: DECREASED
  cellular_components:
  - preferred_term: Microvillus
    term:
      id: GO:0005902
      label: microvillus
  evidence:
  - reference: PMID:18724368
    reference_title: "MYO5B mutations cause microvillus inclusion disease and disrupt epithelial cell polarity."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      MVID is characterized by lack of microvilli on the surface of enterocytes
      and occurrence of intracellular vacuolar structures containing microvilli.
    explanation: >-
      States both halves of this node: absence of surface microvilli and the
      intracellular microvillus-containing vacuoles that define the disease
      histologically.
  downstream:
  - target: Loss of Apical Absorptive and Ion-Transport Capacity
    causal_link_type: DIRECT
- name: Loss of Apical Absorptive and Ion-Transport Capacity
  conforms_to: "enterocyte_polarity_trafficking_failure#Loss of a Functionally Polarized Absorptive Epithelial Surface"
  description: >-
    The functional consequence of losing the apical membrane. The brush border
    is the entire absorptive interface of the enterocyte, carrying the
    disaccharidases, the nutrient carriers and the sodium-coupled ion exchangers
    that drive net fluid absorption. Its loss, together with failure to deliver
    and retain apical transporters, abolishes absorptive capacity across the
    board rather than for one substrate. The resulting diarrhea is therefore
    non-selective and, distinctively, persists during complete bowel rest -
    fluid loss continues when nothing is being fed, because the defect is in the
    absorptive surface itself and not in the handling of any ingested nutrient.
  role: effector
  biological_scale: CELLULAR
  cell_types:
  - preferred_term: Small-intestinal enterocyte
    term:
      id: CL:0000584
      label: enterocyte
  biological_processes:
  - preferred_term: Intestinal absorption
    term:
      id: GO:0050892
      label: intestinal absorption
    modifier: DECREASED
  evidence:
  - reference: PMID:29654747
    reference_title: "Advances in Evaluation of Chronic Diarrhea in Infants."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: >-
      diminution of brush-border microvillar markers, including ezrin, villin,
      phospho-ERM, NHE3, and DPP-IV
    explanation: >-
      Names the specific apical machinery lost at this node, including the
      sodium/hydrogen exchanger NHE3 and the brush-border peptidase DPP-IV -
      direct evidence that losing the surface removes transport and digestive
      capacity together rather than one selective step. Evidence source is OTHER
      because this is a review.
  - reference: PMID:33374714
    reference_title: "Congenital Tufting Enteropathy: Biology, Pathogenesis and Mechanisms."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: >-
      Along with microvillus inclusion disease, a severe disorder belonging to
      the defective enterocyte trafficking and polarity category of CODEs is
      congenital tufting enteropathy (CTE).
    explanation: >-
      Supporting context only: places MVID in the defective enterocyte
      trafficking and polarity CODE class. It establishes the classification, not
      the absorptive-capacity claim, which the preceding citation carries.
  downstream:
  - target: Intractable Secretory Diarrhea and Lifelong Intestinal Failure
    causal_link_type: DIRECT
- name: Intractable Secretory Diarrhea and Lifelong Intestinal Failure
  conforms_to: "enterocyte_polarity_trafficking_failure#Intestinal Failure and Parenteral Nutrition Dependence"
  description: >-
    The clinical output. Profuse watery diarrhea of very high volume begins in
    the first days of life in the early-onset form, causing severe dehydration,
    hyponatremia and metabolic acidosis that require immediate intravenous
    replacement. Because enteral feeding cannot sustain the child, total
    parenteral nutrition is required from the neonatal period and, unlike in
    most causes of intestinal failure, enteral autonomy is essentially never
    achieved. The disease course is then dominated by the complications of
    lifelong parenteral nutrition - intestinal-failure-associated liver disease,
    catheter-related sepsis, growth failure, developmental delay, renal
    impairment and metabolic bone disease.
  role: consequence
  biological_scale: ORGANISM
  locations:
  - preferred_term: Small intestine
    term:
      id: UBERON:0002108
      label: small intestine
  evidence:
  - reference: PMID:21407114
    reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      All of the patients completely depended on PN.
    explanation: >-
      Documents the absolute and universal parenteral-nutrition dependence that
      defines the clinical output of this node in a 24-patient cohort.
  - reference: PMID:21407114
    reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Before or without SBTx, growth failure was common (mean height -2.5
      standard deviations [SD]), as was developmental delay (12/24), liver (20/22
      with fibrosis) or kidney disease (3/23 with moderate renal insufficiency),
      and osteoporosis (6/24).
    explanation: >-
      Quantifies the parenteral-nutrition-dependent complication burden - growth
      failure, developmental delay, liver fibrosis, renal impairment and
      osteoporosis - that dominates the natural history.
- name: Canalicular Trafficking Failure and Low-GGT Intrahepatic Cholestasis
  description: >-
    The hepatic arm of the same trafficking lesion. Myosin Vb organizes apical
    membrane traffic in the hepatocyte just as it does in the enterocyte, and
    the hepatocyte's apical membrane is the bile canaliculus. Loss of the motor
    mislocalizes the canalicular bile-salt export pump and disorganizes the
    canaliculus, producing a low-gamma-glutamyl-transferase intrahepatic
    cholestasis (PFIC10). This arm can occur together with the enteropathy or,
    with certain MYO5B genotypes, in isolation without any intestinal disease,
    which is why MYO5B disease is best understood as one trafficking defect
    expressed in two polarized epithelia. It is mechanistically distinct from -
    and must not be conflated with - the intestinal-failure-associated liver
    disease that parenteral nutrition itself causes in these same children.
  role: effector
  biological_scale: CELLULAR
  locations:
  - preferred_term: Liver
    term:
      id: UBERON:0002107
      label: liver
  evidence:
  - reference: PMID:18724368
    reference_title: "MYO5B mutations cause microvillus inclusion disease and disrupt epithelial cell polarity."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      MYO5B deficiency causes defective trafficking of apical and basolateral
      proteins
    explanation: >-
      Supports the shared trafficking mechanism underlying this arm, which in the
      hepatocyte applies to the canalicular (apical) membrane. This citation
      establishes the trafficking defect, not the hepatic phenotype itself, which
      is evidenced separately below.
  - reference: PMID:27532546
    reference_title: "MYO5B mutations cause cholestasis with normal serum gamma-glutamyl transferase activity in children without microvillous inclusion disease."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Some patients with microvillus inclusion disease due to myosin 5B (MYO5B)
      mutations may develop cholestasis characterized by a progressive familial
      intrahepatic cholestasis-like phenotype with normal serum gamma-glutamyl
      transferase activity.
    explanation: >-
      Establishes the hepatic phenotype of this node in patients who also have
      MVID - a low-GGT, PFIC-like intrahepatic cholestasis attributable to MYO5B
      deficiency.
  - reference: PMID:27532546
    reference_title: "MYO5B mutations cause cholestasis with normal serum gamma-glutamyl transferase activity in children without microvillous inclusion disease."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      These data show that MYO5B deficiency may lead to isolated cholestasis and
      that MYO5B should be considered as an additional progressive familial
      intrahepatic cholestasis gene.
    explanation: >-
      Supports the dissociation claim central to this node: MYO5B deficiency can
      produce cholestasis in the absence of any intestinal disease, which is why
      the hepatic arm is modelled as a parallel effector of the same trafficking
      lesion rather than as a complication of the enteropathy.
  notes: >-
    The specific mechanism by which canalicular trafficking failure produces
    cholestasis - mislocalization of the bile-salt export pump and
    disorganization of the canaliculus - is described in the MYO5B literature but
    is not itself snippet-evidenced here; the cited references establish the
    phenotype and its independence from intestinal disease, not the
    BSEP-mislocalization step. Do not cite the MVID gene-discovery paper
    (PMID:18724368) as evidence for the cholestasis phenotype.
phenotypes:
- name: Intractable Watery Diarrhea
  category: Gastrointestinal
  description: >-
    Profuse, high-volume watery diarrhea beginning in the first days of life
    (early-onset form) or within the first months (late-onset form). It is
    characteristically unresponsive to bowel rest, which distinguishes it from
    the substrate-dependent diet-induced congenital diarrheas.
  phenotype_term:
    preferred_term: Intractable watery diarrhea
    term:
      id: HP:0002014
      label: Diarrhea
    temporality: CHRONIC
  frequency: OBLIGATE
  evidence:
  - reference: PMID:21407114
    reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Microvillous inclusion disease (MVID) is a rare congenital enterocyte
      disorder causing severe diarrhea and intestinal failure.
    explanation: >-
      States severe diarrhea as the defining clinical feature of the disorder.
- name: Villous Atrophy
  category: Histopathological
  description: >-
    Small-intestinal biopsies show villous atrophy with sparse or absent surface
    microvilli, subapical periodic-acid-Schiff-positive granules and the
    pathognomonic intracytoplasmic microvillus inclusions on electron microscopy.
  phenotype_term:
    preferred_term: Villous atrophy
    term:
      id: HP:0011473
      label: Villous atrophy
  evidence:
  - reference: PMID:29654747
    reference_title: "Advances in Evaluation of Chronic Diarrhea in Infants."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: >-
      Other pathological hallmarks of the disease include villus atrophy,
      abnormal subapical periodic acid-Schiff/CD10 staining in enterocytes,
      inclusion staining with antibodies against ezrin or villin
    explanation: >-
      States villus atrophy directly, together with the subapical
      periodic-acid-Schiff/CD10 staining and inclusion staining that accompany it
      - the mucosal architecture claim this phenotype makes, as distinct from the
      microvillar ultrastructure cited on the pathophysiology node.
- name: Dehydration
  category: Metabolic
  description: >-
    Severe dehydration from very high stool water losses, presenting in the
    neonatal period and requiring immediate intravenous fluid and electrolyte
    replacement.
  phenotype_term:
    preferred_term: Dehydration
    term:
      id: HP:0001944
      label: Dehydration
    temporality: ACUTE
  evidence:
  - reference: PMID:29654747
    reference_title: "Advances in Evaluation of Chronic Diarrhea in Infants."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: >-
      Patients with MVID present with severe watery, electrolyte
      transport-related diarrhea and dehydration with failure to thrive beginning
      in the first days or weeks of life.
    explanation: >-
      States dehydration as a presenting feature of MVID specifically, with its
      neonatal onset, rather than as generic guidance about inferring diarrhea.
      Evidence source is OTHER because this is a review.
- name: Failure to Thrive
  category: Growth
  description: >-
    Growth failure is near-universal before transplantation, with a mean height
    of -2.5 standard deviations in a large cohort despite parenteral nutrition.
  phenotype_term:
    preferred_term: Failure to thrive
    term:
      id: HP:0001508
      label: Failure to thrive
    clinical_course: PROGRESSIVE
  frequency: VERY_FREQUENT
  evidence:
  - reference: PMID:21407114
    reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Before or without SBTx, growth failure was common (mean height -2.5
      standard deviations [SD])
    explanation: >-
      Directly quantifies growth failure in the cohort, supporting both the
      phenotype and its VERY_FREQUENT frequency band.
- name: Cholestasis
  category: Hepatic
  description: >-
    Low-gamma-glutamyl-transferase intrahepatic cholestasis from the hepatic
    (canalicular) arm of the MYO5B trafficking defect, which may occur with or
    without the enteropathy. This is distinct from the
    intestinal-failure-associated liver disease caused by long-term parenteral
    nutrition, which affects the great majority of these children independently.
  phenotype_term:
    preferred_term: Cholestasis
    term:
      id: HP:0001396
      label: Cholestasis
  notes: >-
    Curated from clinical description of MYO5B-related disease. Liver fibrosis in
    the cited MVID cohort (20/22) is attributable to parenteral nutrition rather
    than to the MYO5B canalicular defect, so that cohort figure is deliberately
    not used as evidence for this phenotype and no frequency band is asserted.
diagnosis:
- name: Small-intestinal biopsy with electron microscopy
  description: >-
    Duodenal or jejunal biopsy is the diagnostic cornerstone. Light microscopy
    shows villous atrophy without significant inflammatory infiltrate; periodic
    acid-Schiff and CD10 staining reveal abnormal subapical granular deposits;
    electron microscopy demonstrates the pathognomonic intracytoplasmic
    microvillus inclusions and the sparse or absent apical brush border.
  evidence:
  - reference: PMID:18724368
    reference_title: "MYO5B mutations cause microvillus inclusion disease and disrupt epithelial cell polarity."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      MVID is characterized by lack of microvilli on the surface of enterocytes
      and occurrence of intracellular vacuolar structures containing microvilli.
    explanation: >-
      Describes the two ultrastructural findings that establish the diagnosis on
      electron microscopy.
- name: MYO5B and STX3 sequencing
  description: >-
    Molecular confirmation by sequencing of MYO5B and, where MYO5B is negative,
    STX3. Whole-exome or whole-genome sequencing early in the evaluation of
    neonatal intractable diarrhea shortens the diagnostic odyssey and can
    establish the diagnosis before mucosal findings are available.
  evidence:
  - reference: PMID:29654747
    reference_title: "Advances in Evaluation of Chronic Diarrhea in Infants."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: >-
      Whole-exome and genome sequencing in the early stages of evaluation can
      reduce the time required for a definitive diagnosis of CODEs
    explanation: >-
      Supports early genomic testing as the recommended diagnostic strategy for
      congenital diarrheas including MVID. Evidence source is OTHER because this
      is a review.
treatments:
- name: Total Parenteral Nutrition
  description: >-
    Lifelong total parenteral nutrition is required from the neonatal period,
    because the absorptive surface is absent and enteral autonomy is essentially
    never achieved. It is supportive, not disease-modifying: it sustains the
    child while the underlying trafficking defect persists, and it carries its
    own substantial morbidity (intestinal-failure-associated liver disease,
    catheter sepsis, metabolic bone disease, renal impairment). Management in an
    expert intestinal-failure centre measurably improves outcome.
  therapeutic_modality: OTHER
  treatment_term:
    preferred_term: Total Parenteral Nutrition
    term:
      id: NCIT:C29484
      label: Total Parenteral Nutrition
  target_mechanisms:
  - target: Intractable Secretory Diarrhea and Lifelong Intestinal Failure
    treatment_effect: BYPASSES
    description: >-
      Parenteral nutrition bypasses the non-functional intestinal absorptive
      surface entirely, delivering nutrition intravenously; it does not act on
      the trafficking defect or restore the brush border.
    evidence:
    - reference: PMID:21407114
      reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
      supports: SUPPORT
      evidence_source: HUMAN_CLINICAL
      snippet: >-
        PN in MVID is difficult to manage and requires expertise.
      explanation: >-
        Supports parenteral nutrition as the mainstay of management and records
        that outcome depends on the expertise with which it is delivered.
- name: Small-Bowel Transplantation
  description: >-
    Small-bowel transplantation, isolated or combined with liver transplantation
    when intestinal-failure-associated liver disease has advanced, is the only
    definitive treatment. It replaces the defective epithelium with donor
    intestine carrying two functional MYO5B alleles. In a 24-patient cohort,
    survival was 77% with transplantation versus 63% without, and catch-up growth
    followed transplantation in some children.
  therapeutic_modality: SURGERY
  treatment_term:
    preferred_term: Small Bowel Transplantation
    term:
      id: NCIT:C157985
      label: Small Bowel Transplantation
  target_mechanisms:
  - target: Loss of Apical Absorptive and Ion-Transport Capacity
    treatment_effect: RESTORES
    description: >-
      Replacing the intestine with genotypically normal donor epithelium restores
      an intact brush border and absorptive capacity, the only intervention that
      addresses the effector node rather than compensating for it.
    evidence:
    - reference: PMID:21407114
      reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
      supports: SUPPORT
      evidence_source: HUMAN_CLINICAL
      snippet: >-
        SBTx, despite being complicated, remains the only hope to improve the
        quality of life and long-term prognosis of these children.
      explanation: >-
        States that small-bowel transplantation is the only intervention that
        improves long-term prognosis, supporting it as the definitive treatment.
    - reference: PMID:21407114
      reference_title: "Microvillous inclusion disease: how to improve the prognosis of a severe congenital enterocyte disorder."
      supports: SUPPORT
      evidence_source: HUMAN_CLINICAL
      snippet: >-
        Patient survival rates were 63% without SBTx and 77% with SBTx.
      explanation: >-
        Quantifies the survival difference associated with transplantation in the
        cohort.
notes: >-
  MVID sits in the "defective enterocyte trafficking and polarity" class of
  congenital diarrheas and enteropathies, alongside congenital tufting
  enteropathy. It deliberately does NOT conform to the
  diet_induced_osmotic_diarrhea module: that module requires a normal
  villus-to-crypt ratio and substrate-dependent diarrhea that remits on dietary
  withdrawal, whereas MVID has villous atrophy and diarrhea that persists during
  complete bowel rest. The two disorders are distinguished at exactly this point
  in the current CODE nosology (PMID:29654747). MVID is curated as a conformer of the
  enterocyte_polarity_trafficking_failure module, which factors out the chain it
  shares with congenital tufting enteropathy. That module admits two molecularly
  distinct trigger arms - apical-delivery trafficking failure (this entry, via
  MYO5B/STX3) and junctional adhesion failure (tufting enteropathy, via
  EpCAM) - which converge on loss of a functionally polarized absorptive
  surface. The arms are not equivalent and their histologies differ, so this
  entry substitutes its own lesion rather than the module's generic description.