Hypertrophic Cardiomyopathy 1

Genetic MONDO:0008647 Pathograph 9 Show in embeddings browser Hypertrophic Cardiomyopathy Genetic Disorder

Hypertrophic cardiomyopathy 1 (CMH1) is the MYH7-related form of familial hypertrophic cardiomyopathy and, with the MYBPC3 form (CMH4), one of the two commonest genetic causes of the disease. MYH7 encodes the beta-cardiac myosin heavy chain (beta-MyHC), the force-generating molecular motor of the cardiac thick filament. CMH1 is historically the founding molecular diagnosis of the entire field: the first mutation ever shown to cause familial hypertrophic cardiomyopathy was the MYH7 Arg403Gln (R403Q) missense change, identified in a large kindred in 1990. Unlike MYBPC3-related disease, where truncating alleles act through haploinsufficiency, the great majority of MYH7 disease alleles are missense variants that are stably expressed and incorporated into the sarcomere as a mutant motor. The operative mechanism is therefore not loss of protein but a gain of contractile function: the mutant myosin destabilizes the folded-back, autoinhibited super-relaxed state of the thick filament, freeing more heads to interact with actin, and often raises the intrinsic force and the energetic cost of each cross-bridge. The net cellular consequence is hypercontractility with impaired relaxation and raised tension cost, which drives the structural remodeling — cardiomyocyte hypertrophy, myofiber disarray, and interstitial fibrosis — that defines the clinical phenotype. Inheritance is autosomal dominant with incomplete, age-related penetrance; MYH7 variants carry a clinically important risk of malignant ventricular arrhythmia and sudden cardiac death, and beta-cardiac myosin is the direct molecular target of the cardiac myosin inhibitors (mavacamten, aficamten) now used to treat obstructive disease.

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1
Inheritance
7
Pathophys.
7
Phenotypes
9
Pathograph
1
Genes
3
Medical Actions
1
Trials
1
References
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Classifications

Harrison's Part
CARDIOVASCULAR GENETICS ENVIRONMENT DISEASE
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Inheritance

1
Autosomal dominant HP:0000006
CMH1 is inherited as an autosomal dominant trait, with a single mutant MYH7 allele producing disease. Penetrance is incomplete and age-related, and a family history of hypertrophic cardiomyopathy and of sudden cardiac death is characteristic of sarcomere-variant-positive disease, so cascade screening of relatives must be longitudinal rather than a single evaluation.
Autosomal dominant inheritance Penetrance: INCOMPLETE
Show evidence (2 references)
"MYH7 | HGNC:7577 | hypertrophic cardiomyopathy | MONDO:0005045 | AD | Definitive"
ClinGen's Hereditary Cardiovascular Disease Gene Curation Expert Panel classifies the MYH7-hypertrophic cardiomyopathy relationship as Definitive with autosomal dominant inheritance.
PMID:23674365 SUPPORT Human Clinical
"The presence of any sarcomere gene mutation was associated with a younger age at presentation (38.4 vs 46.0 years, p<0.0005), a family history of HCM (50.6% vs 23.1%, p<0.0005), a family history of SCD (27.0% vs 14.9%, p<0.0005) and greater MLVWT (21.0 vs 19.3 mm, p=0.03)."
Meta-analysis of 2459 patients documenting the familial, dominantly inherited pattern of sarcomere-positive HCM, with a positive family history of both HCM and sudden cardiac death and younger onset than sarcomere-negative disease.

Pathophysiology

7
MYH7 Missense Variant in Beta-Cardiac Myosin Heavy Chain
The initiating lesion in CMH1 is a heterozygous missense variant in MYH7, the gene encoding the beta-cardiac myosin heavy chain — the ATP-hydrolysing motor of the sarcomeric thick filament. The founding example, and still the archetype, is Arg403Gln (R403Q), the first mutation ever linked to familial hypertrophic cardiomyopathy; R403Q sits in the globular myosin head (S1), the region responsible for actin interaction and motor function. In contrast to the truncating, haploinsufficiency mechanism of MYBPC3 disease, most MYH7 alleles are missense changes that are stably expressed and incorporated into the sarcomere as a mutant motor, so the mutant protein perturbs contraction directly rather than by its absence.
Cardiomyocyte CL:0000746 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiomyocyte, annotated with cardiac muscle cell (CL:0000746). CL:0000746 is a cell type from the Cell Ontology.
MYH7 hgnc:7577 HUGO Gene Nomenclature Committee (hgnc) Relation: this pathophysiological event involves this gene This pathophysiological event involves MYH7 (hgnc:7577). hgnc:7577 is a gene from the HUGO Gene Nomenclature Committee.
Sarcomere Organization GO:0045214 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Sarcomere Organization (GO:0045214). GO:0045214 is a biological process from the Gene Ontology. ⚠ ABNORMAL
Myosin thick filament GO:0032982 Gene Ontology (GO) Relation: this pathophysiological event involves this cellular component This pathophysiological event involves Myosin thick filament, annotated with myosin filament (GO:0032982). GO:0032982 is a cellular component from the Gene Ontology.
Show evidence (2 references)
PMID:1975517 SUPPORT Human Clinical
"A point mutation in exon 13 of the beta cardiac myosin heavy chain (MHC) gene is present in all individuals affected with familial hypertrophic cardiomyopathy (FHC) from a large kindred. This missense mutation converts a highly conserved arginine residue (Arg-403) to a glutamine."
The founding CMH1 report: co-segregation of the MYH7 R403Q missense variant with familial hypertrophic cardiomyopathy in a large kindred, establishing the beta-cardiac myosin heavy chain gene as a cause of the disease.
PMID:23798412 SUPPORT In Vitro
"HCM is caused by mutations in the genes encoding the fundamental force-generating machinery of the cardiac muscle, including β-cardiac myosin."
Places the MYH7 lesion in the force-generating machinery of the sarcomere, the trigger for this entity.
Destabilization of the Autoinhibited Super-Relaxed State
In the resting sarcomere a large fraction of myosin heads occupy the super-relaxed (SRX) state — folded back against the thick-filament backbone in the interacting-heads motif (IHM), with very low ATPase activity, forming an energy-conserving reserve of motors withheld from force generation. Many MYH7 HCM mutations map to the interfaces that stabilize this folded-back state (the myosin mesa, the head-head and head-tail contacts), and they weaken those intramolecular interactions. The result is release of sequestered heads from the SRX/IHM into the disordered-relaxed state, so more heads become functionally accessible to actin — the molecular origin of hypercontractility shared across a large subset of MYH7 alleles.
Cardiomyocyte CL:0000746 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiomyocyte, annotated with cardiac muscle cell (CL:0000746). CL:0000746 is a cell type from the Cell Ontology.
ATP Hydrolysis by Myosin GO:0016887 Gene Ontology (GO) Relation: this pathophysiological event involves this molecular function This pathophysiological event involves increased ATP Hydrolysis by Myosin, annotated with ATP hydrolysis activity (GO:0016887). GO:0016887 is a molecular function from the Gene Ontology. ↑ INCREASED
Myosin thick filament GO:0032982 Gene Ontology (GO) Relation: this pathophysiological event involves this cellular component This pathophysiological event involves Myosin thick filament, annotated with myosin filament (GO:0032982). GO:0032982 is a cellular component from the Gene Ontology.
Show evidence (4 references)
PMID:31213605 SUPPORT In Vitro
"folded-back sequestered state referred to as the interacting heads motif (IHM) lead to a significant increase in the number of heads functionally accessible for interaction with actin."
Direct biochemical evidence that MYH7 HCM mutations at IHM interfaces release sequestered heads, increasing the number available to interact with actin.
PMID:28481356 SUPPORT In Vitro
"hypercontractility is due to an increase in the number of myosin heads (S1) that are accessible for force production."
States the core hypothesis this node captures: HCM hypercontractility arises from an increase in the number of force-producing heads freed from the sequestered state.
PMID:34117120 SUPPORT In Vitro
"destabilized the super relaxed state in longer, two-headed myosin constructs, freeing more heads to generate force."
Shows a specific MYH7 mutation (P710R) destabilizing the super-relaxed state and freeing heads for force generation.
+ 1 more reference
Increased Motor Force and Energetic Cost
Beyond releasing sequestered heads, individual MYH7 missense alleles alter the mechanochemistry of the motor stroke itself. Single-molecule and myofibril studies show mutation-specific changes — increased intrinsic force per cross-bridge, and faster cross-bridge detachment kinetics — that together raise the force generated by the ensemble of motors and increase the ATP consumed per unit tension (tension cost). The energetic penalty, measured directly in human myocardium carrying R403Q, is a distinctive feature of the beta-myosin motor lesion and links the molecular defect to the myocardial energy deficit seen in HCM.
Cardiomyocyte CL:0000746 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiomyocyte, annotated with cardiac muscle cell (CL:0000746). CL:0000746 is a cell type from the Cell Ontology.
Actin-Myosin Filament Sliding GO:0033275 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Actin-Myosin Filament Sliding (GO:0033275). GO:0033275 is a biological process from the Gene Ontology. ⚠ ABNORMAL
ATP Hydrolysis by Myosin GO:0016887 Gene Ontology (GO) Relation: this pathophysiological event involves this molecular function This pathophysiological event involves increased ATP Hydrolysis by Myosin, annotated with ATP hydrolysis activity (GO:0016887). GO:0016887 is a molecular function from the Gene Ontology. ↑ INCREASED
Show evidence (3 references)
PMID:23798412 SUPPORT In Vitro
"The major change in the R453C human β-cardiac subfragment 1 is a 50% increase in the intrinsic force of the motor compared with wild type, with no appreciable change in the stroke size, as observed with a dual-beam optical trap."
Single-molecule optical-trap measurement of a raised intrinsic force for an MYH7 mutant motor, predicting a hypercontractile ensemble.
PMID:23798412 SUPPORT In Vitro
"Overall, this study suggests that the R453C mutation should result in a hypercontractile state in the heart muscle."
States the functional conclusion — a hypercontractile state — that the motor-level force change produces.
PMID:24928957 SUPPORT Human Clinical
"This correlation suggests that faster cross-bridge relaxation kinetics results in an increase in energetic cost of tension generation in human HCM with the R403Q mutation compared to HCMsmn."
Direct measurement in human R403Q myocardium (single myofibrils and muscle strips) linking faster cross-bridge kinetics to an increased energetic cost of tension generation, evidence obtained in patient cardiac tissue.
Sarcomere Hypercontractility with Impaired Relaxation
At the cell level the molecular changes converge on hypercontractility: CRISPR-edited human iPSC-derived cardiomyocytes carrying an MYH7 HCM mutation generate significantly greater contractile force than isogenic controls. Because the same lesion disturbs cross-bridge relaxation kinetics, systolic hypercontraction is accompanied by impaired diastolic relaxation. This hypercontractile, poorly relaxing cell state — a key pathophysiological abnormality of HCM and the determinant of dynamic outflow obstruction — is the cellular phenotype the cardiac myosin inhibitors were designed to normalize.
Ventricular cardiomyocyte CL:0002131 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Ventricular cardiomyocyte, annotated with regular ventricular cardiac myocyte (CL:0002131). CL:0002131 is a cell type from the Cell Ontology.
Cardiac Muscle Cell Contraction GO:0086003 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves increased Cardiac Muscle Cell Contraction (GO:0086003). GO:0086003 is a biological process from the Gene Ontology. ↑ INCREASED Regulation of Cardiac Muscle Contraction GO:0055117 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Regulation of Cardiac Muscle Contraction (GO:0055117). GO:0055117 is a biological process from the Gene Ontology. ⚠ ABNORMAL
Show evidence (2 references)
PMID:34117120 SUPPORT In Vitro
"produced significantly increased force (measured by traction force microscopy) compared with isogenic control cells."
Isogenic human iPSC-cardiomyocyte evidence that an MYH7 HCM mutation increases cellular contractile force.
PMID:32871100 SUPPORT Human Clinical
"Cardiac muscle hypercontractility is a key pathophysiological abnormality in hypertrophic cardiomyopathy, and a major determinant of dynamic left ventricular outflow tract (LVOT) obstruction."
Confirms hypercontractility as the central pathophysiological abnormality of HCM and links it to outflow obstruction; PARTIAL because the trial enrolled obstructive HCM broadly rather than genotyped MYH7 carriers.
Cell-to-Cell Contractile Imbalance
A mechanism specific to the missense/poison-peptide biology of MYH7 disease. Because the mutant and wild-type MYH7 alleles are transcribed in a random, burst-like fashion and independently of one another, the fraction of mutant beta-myosin varies markedly from cardiomyocyte to cardiomyocyte within a single patient's myocardium. This produces much greater cell-to-cell variability of contractile function than in control hearts. The resulting mechanical mismatch between adjacent, differently loaded myocytes is proposed to drive the myofiber disarray and interstitial fibrosis that are hallmarks of HCM, providing a route from the molecular lesion to the tissue phenotype that does not depend on the average level of hypercontractility alone.
Cardiomyocyte CL:0000746 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiomyocyte, annotated with cardiac muscle cell (CL:0000746). CL:0000746 is a cell type from the Cell Ontology.
Cardiac Muscle Cell Contraction GO:0086003 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Cardiac Muscle Cell Contraction (GO:0086003). GO:0086003 is a biological process from the Gene Ontology. ⚠ ABNORMAL
Show evidence (2 references)
PMID:30740621 SUPPORT Human Clinical
"substantial contractile variability from cardiomyocyte to cardiomyocyte within a patient's myocardium, much higher than in controls."
Documents the cell-to-cell contractile variability in beta-MyHC-mutant patient myocardium that this node captures.
PMID:30740621 SUPPORT Human Clinical
"contractile imbalance due to unequal fractions of mutated and wildtype protein among individual cardiomyocytes over time will induce cardiomyocyte disarray and fibrosis, hallmarks of HCM."
States the contractile-imbalance hypothesis linking unequal mutant-allele expression to disarray and fibrosis, the hallmarks of HCM.
Ventricular Hypertrophy, Myocyte Disarray and Fibrosis
Chronic hypercontractility with raised energetic cost, amplified by cell-to-cell contractile imbalance, drives the structural remodeling that defines the clinical phenotype: cardiomyocyte hypertrophy — typically asymmetric and septal-predominant — together with myofiber disarray and interstitial and replacement fibrosis. Pro-hypertrophic growth signalling contributes: in an MYH7-mutant human iPSC-cardiomyocyte model, cellular hypertrophy was prevented by inhibition of ERK or Akt, implicating those pathways downstream of the mechanical stimulus. Fibrosis provides the substrate for both diastolic dysfunction and re-entrant arrhythmia.
Cardiomyocyte CL:0000746 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiomyocyte, annotated with cardiac muscle cell (CL:0000746). CL:0000746 is a cell type from the Cell Ontology. Cardiac Fibroblast CL:0002548 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiac Fibroblast, annotated with fibroblast of cardiac tissue (CL:0002548). CL:0002548 is a cell type from the Cell Ontology.
Cardiac Muscle Hypertrophy in Response to Stress GO:0014898 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves increased Cardiac Muscle Hypertrophy in Response to Stress (GO:0014898). GO:0014898 is a biological process from the Gene Ontology. ↑ INCREASED Extracellular Matrix Organization GO:0030198 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves increased Extracellular Matrix Organization (GO:0030198). GO:0030198 is a biological process from the Gene Ontology. ↑ INCREASED
Interventricular septum UBERON:0002094 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Interventricular septum (UBERON:0002094). UBERON:0002094 is an anatomical location from the Uberon multi-species anatomy ontology. Left ventricle UBERON:0002084 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Left ventricle, annotated with heart left ventricle (UBERON:0002084). UBERON:0002084 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (3 references)
PMID:28912181 SUPPORT Human Clinical
"The histological features of HCM include myocyte hypertrophy and disarray, as well as interstitial fibrosis."
Names the defining tissue-level triad — myocyte hypertrophy, disarray, and interstitial fibrosis — that this node represents.
PMID:34117120 SUPPORT In Vitro
"Cellular hypertrophy was prevented in the P710R cells by inhibition of ERK or Akt."
Implicates ERK and Akt growth signalling downstream of the MYH7 mechanical lesion in producing the cellular hypertrophy of this node.
PMID:30740621 SUPPORT Human Clinical
"contractile imbalance due to unequal fractions of mutated and wildtype protein among individual cardiomyocytes over time will induce cardiomyocyte disarray and fibrosis, hallmarks of HCM."
Ties the upstream contractile imbalance to the disarray and fibrosis of this tissue node.
Diastolic Dysfunction, Outflow Obstruction, Heart Failure and Arrhythmic Risk
The hypertrophied, disarrayed and fibrotic ventricle produces the clinical endpoints of CMH1. Hypercontractility with septal hypertrophy generates dynamic left ventricular outflow tract obstruction in a subset of patients; the stiff, poorly relaxing ventricle causes diastolic dysfunction; and the fibrotic, electrically heterogeneous substrate carries a risk of malignant ventricular arrhythmia and sudden cardiac death that is clinically important for MYH7 variants. Sarcomere-positive disease, of which MYH7 is a leading cause, presents younger and with a stronger family history of sudden death than sarcomere-negative HCM.
Cardiomyocyte CL:0000746 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Cardiomyocyte, annotated with cardiac muscle cell (CL:0000746). CL:0000746 is a cell type from the Cell Ontology.
Heart Contraction GO:0060047 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Heart Contraction (GO:0060047). GO:0060047 is a biological process from the Gene Ontology. ⚠ ABNORMAL
Show evidence (2 references)
PMID:23674365 SUPPORT Human Clinical
"There were no differences when the two most frequently affected genes, MYBPC3 and MYH7, were compared."
Records the important nuance that, at the cohort level, MYH7 and MYBPC3 do not differ significantly across the pooled clinical features, so MYH7 is not simply a uniformly more severe genotype than MYBPC3; it is one of the two commonest sarcomeric causes.
PMID:39681440 SUPPORT Human Clinical
"In the five families with HCM carrying MYH7-R453C mutations, genotype is highly correlated with clinical phenotype, and patients have a high risk of sudden death and poor prognosis."
A family study of a specific MYH7 variant documenting high sudden-death risk and poor prognosis, the arrhythmic endpoint of this node.

Pathograph

Use the checkboxes to hide or show graph categories. Hover nodes for evidence and cross-linked metadata.
Pathograph: causal mechanism network for Hypertrophic Cardiomyopathy 1 Interactive directed graph showing how pathophysiology mechanisms, phenotypes, genetic factors and variants, experimental models, environmental triggers, and treatments relate through causal and linked edges.

Phenotypes

7
Cardiovascular 5
Hypertrophic Cardiomyopathy OBLIGATE HP:0001639 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Hypertrophic cardiomyopathy (HP:0001639). HP:0001639 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:28912181 SUPPORT Human Clinical
"Hypertrophic cardiomyopathy (HCM) is a genetic disorder that is characterized by left ventricular hypertrophy unexplained by secondary causes and a nondilated left ventricle with preserved or increased ejection fraction."
States the defining clinical feature — unexplained left ventricular hypertrophy with a nondilated ventricle — of the disease this MYH7 form manifests.
Left Ventricular Diastolic Dysfunction HP:0025168 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Left ventricular diastolic dysfunction (HP:0025168). HP:0025168 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:24928957 SUPPORT Human Clinical
"Cross-bridge slow relaxation kinetics in single R403Q myofibrils was significantly higher (P < 0.0001) than in HCMsmn myofibrils"
Documents disturbed cross-bridge relaxation kinetics in human R403Q myocardium, the molecular basis of the impaired relaxation underlying diastolic dysfunction.
Sudden Cardiac Death HP:0001645 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Sudden cardiac death (HP:0001645). HP:0001645 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:39681440 SUPPORT Human Clinical
"In the five families with HCM carrying MYH7-R453C mutations, genotype is highly correlated with clinical phenotype, and patients have a high risk of sudden death and poor prognosis."
A family study of an MYH7 variant documenting a high risk of sudden death.
Ventricular Arrhythmia HP:0004308 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Ventricular arrhythmia (HP:0004308). HP:0004308 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:39681440 SUPPORT Human Clinical
"2 experienced (F1Ⅲ1, F3Ⅲ3) events of sudden cardiac death survival"
Aborted sudden cardiac death (survived cardiac arrest) in MYH7-R453C carriers evidences the malignant ventricular arrhythmia of this node.
Congestive Heart Failure HP:0001635 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Congestive heart failure (HP:0001635). HP:0001635 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:39681440 SUPPORT Human Clinical
"2(F1Ⅱ2, F3Ⅱ1) died from heart failure during the follow-up period."
Deaths from heart failure among MYH7-R453C carriers evidence heart failure as a clinical endpoint of the disease.
Other 2
Asymmetric Septal Hypertrophy HP:0001670 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Asymmetric septal hypertrophy (HP:0001670). HP:0001670 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:23674365 SUPPORT Human Clinical
"The presence of any sarcomere gene mutation was associated with a younger age at presentation (38.4 vs 46.0 years, p<0.0005), a family history of HCM (50.6% vs 23.1%, p<0.0005), a family history of SCD (27.0% vs 14.9%, p<0.0005) and greater MLVWT (21.0 vs 19.3 mm, p=0.03)."
Sarcomere-positive HCM, of which MYH7 is a leading cause, shows greater maximum left ventricular wall thickness than sarcomere-negative disease, supporting the pronounced hypertrophy of this member.
Left Ventricular Outflow Tract Obstruction HP:0032092 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Left ventricular outflow tract obstruction (HP:0032092). HP:0032092 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:32871100 SUPPORT Human Clinical
"Cardiac muscle hypercontractility is a key pathophysiological abnormality in hypertrophic cardiomyopathy, and a major determinant of dynamic left ventricular outflow tract (LVOT) obstruction."
Links hypercontractility to dynamic LVOT obstruction; PARTIAL because the trial population is obstructive HCM broadly rather than genotyped MYH7 carriers.
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Genetic Associations

1
MYH7
Gene: MYH7 hgnc:7577 HUGO Gene Nomenclature Committee (hgnc) Relation: this disease-associated gene is this gene This disease-associated gene is MYH7 (hgnc:7577). hgnc:7577 is a gene from the HUGO Gene Nomenclature Committee. relationship_type: CAUSATIVE
Show evidence (2 references)
"MYH7 | HGNC:7577 | hypertrophic cardiomyopathy | MONDO:0005045 | AD | Definitive"
ClinGen's Gene Curation Expert Panel classifies this gene-disease relationship as Definitive, which is the authority for curating the gene as CAUSATIVE rather than a candidate.
PMID:23674365 SUPPORT Human Clinical
"The presence of any sarcomere gene mutation was associated with a younger age at presentation (38.4 vs 46.0 years, p<0.0005), a family history of HCM (50.6% vs 23.1%, p<0.0005), a family history of SCD (27.0% vs 14.9%, p<0.0005) and greater MLVWT (21.0 vs 19.3 mm, p=0.03)."
The cohort is sarcomere-mutation carriers across genes rather than MYH7 carriers, so this is class-level evidence that a sarcomere genotype shifts age at presentation and family history. Curated PARTIAL for that reason; the same study's finding of no outcome difference between MYH7 and MYBPC3 carriers is recorded in this entry's pathophysiology.
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Medical Actions

3
Mavacamten (Cardiac Myosin Inhibitor)
Action: PharmacotherapyNCI Thesaurus (NCIT) Relation: this treatment is this clinical intervention This treatment is Pharmacotherapy (NCIT:C15986). NCIT:C15986 is a clinical intervention from the NCI Thesaurus. NCIT:C15986
Agent: mavacamten CHEBI:756998 Chemical Entities of Biological Interest (CHEBI) Relation: this treatment uses this therapeutic agent This treatment uses mavacamten (CHEBI:756998). CHEBI:756998 is a therapeutic agent from Chemical Entities of Biological Interest.
Mavacamten is a small-molecule allosteric inhibitor of cardiac myosin ATPase that reduces the number of force-generating actin-myosin cross-bridges, stabilizing myosin in the energy-sparing super-relaxed state. It directly counteracts the hypercontractility that is the central abnormality of HCM, and is approved for symptomatic obstructive hypertrophic cardiomyopathy; because beta-cardiac myosin is the MYH7 gene product, the drug acts on the same motor that carries the CMH1 lesion.
Mechanism Target:
INHIBITS Destabilization of the Autoinhibited Super-Relaxed State — Mavacamten stabilizes the super-relaxed state and reduces the number of heads available for force generation, directly opposing the SRX destabilization that drives hypercontractility.
Show evidence (1 reference)
PMID:40118457 SUPPORT In Vitro
"cardiac myosin inhibitor mavacamten slows nucleotide release by an equal factor for both heavy meromyosin and subfragment 1, thus only indirectly influencing the occupancy time of the SRX state."
Establishes mavacamten's biochemical action on beta-cardiac myosin nucleotide handling and its effect on SRX-state occupancy, the node it targets.
Show evidence (1 reference)
PMID:32871100 SUPPORT Human Clinical
"Cardiac muscle hypercontractility is a key pathophysiological abnormality in hypertrophic cardiomyopathy, and a major determinant of dynamic left ventricular outflow tract (LVOT) obstruction."
The pivotal phase 3 trial establishing the rationale and use of a cardiac myosin inhibitor targeting hypercontractility in obstructive HCM.
Septal Reduction (Surgical Myectomy)
Action: septal myectomyNCI Thesaurus (NCIT) Relation: this treatment is this clinical intervention This treatment is septal myectomy, annotated with Myectomy (NCIT:C51591). NCIT:C51591 is a clinical intervention from the NCI Thesaurus. Ontology label: Myectomy NCIT:C51591
Surgical septal myectomy relieves severe drug-refractory left ventricular outflow tract obstruction by resecting hypertrophied basal septal muscle.
Implantable Cardioverter-Defibrillator
Action: implantable cardioverter-defibrillator placementNCI Thesaurus (NCIT) Relation: this treatment is this clinical intervention This treatment is implantable cardioverter-defibrillator placement (NCIT:C80435). NCIT:C80435 is a clinical intervention from the NCI Thesaurus. Ontology label: Implantable Cardioverter-Defibrillator Placement NCIT:C80435
An implantable cardioverter-defibrillator provides secondary or primary prevention of sudden cardiac death in patients judged to be at high arrhythmic risk, terminating malignant ventricular arrhythmia.
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Prevalence

2
Worldwide (hypertrophic cardiomyopathy overall)
Point Prevalence 200.0 per 100,000 >1 in 1,000
Clinical prevalence of hypertrophic cardiomyopathy in the general population, the context for the MYH7 subtype. MYH7-specific population rates are not separately documented.
Show evidence (1 reference)
PMID:31213605 SUPPORT Other
"Hypertrophic cardiomyopathy (HCM) affects 1 in 500 people and leads to hyper-contractility of the heart."
Gives the ~1 in 500 general-population prevalence of hypertrophic cardiomyopathy overall. Evidence source is OTHER because this is a biochemical mechanism paper stating the epidemiological background.
Worldwide (MYH7 share of HCM mutations)
Unknown Not yet documented
No population-based prevalence estimate exists for the MYH7-specific entity. MYH7 is, with MYBPC3, one of the two most frequently mutated genes in hypertrophic cardiomyopathy, accounting for roughly 40% of identified HCM-causing mutations.
Show evidence (1 reference)
PMID:31213605 SUPPORT Other
"Nearly 40 percent of HCM-causing mutations are found in human β-cardiac myosin."
Quantifies MYH7's share of HCM-causing mutations. Evidence source is OTHER because this is a biochemical mechanism paper.
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Clinical Trials

1
NCT01912534 PHASE_II COMPLETED
VANISH (Valsartan for Attenuating Disease Evolution in Early Sarcomeric Hypertrophic Cardiomyopathy) tested whether the angiotensin receptor blocker valsartan can modify disease progression in young carriers of sarcomeric gene mutations, including MYH7, with early-stage HCM.
Target Phenotypes: Hypertrophic cardiomyopathy HP:0001639 Human Phenotype Ontology (HP) Relation: this clinical trial targets this phenotype This clinical trial targets Hypertrophic cardiomyopathy (HP:0001639). HP:0001639 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:31813281 SUPPORT Human Clinical
"targeted young sarcomeric gene mutation carriers with early-stage hypertrophic cardiomyopathy (HCM) to test whether valsartan can modify disease progression."
Describes the VANISH trial's design and its enrollment of young sarcomeric (including MYH7) mutation carriers with early HCM.
{ }

Source YAML

click to show
name: Hypertrophic Cardiomyopathy 1
creation_date: "2026-08-22T00:00:00Z"
synonyms:
- CMH1
- MYH7 hypertrophic cardiomyopathy
- beta-myosin heavy chain hypertrophic cardiomyopathy
- cardiomyopathy, familial hypertrophic, 1
- familial hypertrophic cardiomyopathy type 1
- hypertrophic cardiomyopathy caused by mutation in MYH7
description: >-
  Hypertrophic cardiomyopathy 1 (CMH1) is the MYH7-related form of familial
  hypertrophic cardiomyopathy and, with the MYBPC3 form (CMH4), one of the two
  commonest genetic causes of the disease. MYH7 encodes the beta-cardiac myosin
  heavy chain (beta-MyHC), the force-generating molecular motor of the cardiac
  thick filament. CMH1 is historically the founding molecular diagnosis of the
  entire field: the first mutation ever shown to cause familial hypertrophic
  cardiomyopathy was the MYH7 Arg403Gln (R403Q) missense change, identified in a
  large kindred in 1990.
  Unlike MYBPC3-related disease, where truncating alleles act through
  haploinsufficiency, the great majority of MYH7 disease alleles are missense
  variants that are stably expressed and incorporated into the sarcomere as a
  mutant motor. The operative mechanism is therefore not loss of protein but a
  gain of contractile function: the mutant myosin destabilizes the folded-back,
  autoinhibited super-relaxed state of the thick filament, freeing more heads to
  interact with actin, and often raises the intrinsic force and the energetic
  cost of each cross-bridge. The net cellular consequence is hypercontractility
  with impaired relaxation and raised tension cost, which drives the structural
  remodeling — cardiomyocyte hypertrophy, myofiber disarray, and interstitial
  fibrosis — that defines the clinical phenotype. Inheritance is autosomal
  dominant with incomplete, age-related penetrance; MYH7 variants carry a
  clinically important risk of malignant ventricular arrhythmia and sudden
  cardiac death, and beta-cardiac myosin is the direct molecular target of the
  cardiac myosin inhibitors (mavacamten, aficamten) now used to treat obstructive
  disease.
category: Genetic
classifications:
  harrisons_chapter:
  - classification_value: CARDIOVASCULAR
  - classification_value: GENETICS_ENVIRONMENT_DISEASE
disease_term:
  preferred_term: hypertrophic cardiomyopathy 1
  term:
    id: MONDO:0008647
    label: hypertrophic cardiomyopathy 1
parents:
- Hypertrophic Cardiomyopathy
- Genetic Disorder
inheritance:
- name: Autosomal dominant
  description: >-
    CMH1 is inherited as an autosomal dominant trait, with a single mutant MYH7
    allele producing disease. Penetrance is incomplete and age-related, and a
    family history of hypertrophic cardiomyopathy and of sudden cardiac death is
    characteristic of sarcomere-variant-positive disease, so cascade screening of
    relatives must be longitudinal rather than a single evaluation.
  inheritance_term:
    preferred_term: Autosomal dominant inheritance
    term:
      id: HP:0000006
      label: Autosomal dominant inheritance
  penetrance: INCOMPLETE
  evidence:
  - reference: CGGV:assertion_31325c90-05cb-4db0-9372-e4f705cd5c82-2023-07-12T160000.000Z
    reference_title: "MYH7 / hypertrophic cardiomyopathy (Definitive)"
    supports: SUPPORT
    evidence_source: OTHER
    snippet: "MYH7 | HGNC:7577 | hypertrophic cardiomyopathy | MONDO:0005045 | AD | Definitive"
    explanation: >-
      ClinGen's Hereditary Cardiovascular Disease Gene Curation Expert Panel
      classifies the MYH7-hypertrophic cardiomyopathy relationship as Definitive
      with autosomal dominant inheritance.
  - reference: PMID:23674365
    reference_title: "A systematic review and meta-analysis of genotype-phenotype associations in patients with hypertrophic cardiomyopathy caused by sarcomeric protein mutations."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      The presence of any sarcomere gene mutation was associated with a younger
      age at presentation (38.4 vs 46.0 years, p<0.0005), a family history of HCM
      (50.6% vs 23.1%, p<0.0005), a family history of SCD (27.0% vs 14.9%,
      p<0.0005) and greater MLVWT (21.0 vs 19.3 mm, p=0.03).
    explanation: >-
      Meta-analysis of 2459 patients documenting the familial, dominantly
      inherited pattern of sarcomere-positive HCM, with a positive family history
      of both HCM and sudden cardiac death and younger onset than
      sarcomere-negative disease.
prevalence:
- population: Worldwide (hypertrophic cardiomyopathy overall)
  measure_type: POINT_PREVALENCE
  prevalence_class: ABOVE_1_IN_1000
  rate_per_100000: 200.0
  notes: >-
    Clinical prevalence of hypertrophic cardiomyopathy in the general population,
    the context for the MYH7 subtype. MYH7-specific population rates are not
    separately documented.
  evidence:
  - reference: PMID:31213605
    reference_title: "β-Cardiac myosin hypertrophic cardiomyopathy mutations release sequestered heads and increase enzymatic activity."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: "Hypertrophic cardiomyopathy (HCM) affects 1 in 500 people and leads to hyper-contractility of the heart."
    explanation: >-
      Gives the ~1 in 500 general-population prevalence of hypertrophic
      cardiomyopathy overall. Evidence source is OTHER because this is a
      biochemical mechanism paper stating the epidemiological background.
- population: Worldwide (MYH7 share of HCM mutations)
  measure_type: UNKNOWN
  prevalence_class: NOT_YET_DOCUMENTED
  notes: >-
    No population-based prevalence estimate exists for the MYH7-specific entity.
    MYH7 is, with MYBPC3, one of the two most frequently mutated genes in
    hypertrophic cardiomyopathy, accounting for roughly 40% of identified
    HCM-causing mutations.
  evidence:
  - reference: PMID:31213605
    reference_title: "β-Cardiac myosin hypertrophic cardiomyopathy mutations release sequestered heads and increase enzymatic activity."
    supports: SUPPORT
    evidence_source: OTHER
    snippet: "Nearly 40 percent of HCM-causing mutations are found in human β-cardiac myosin."
    explanation: >-
      Quantifies MYH7's share of HCM-causing mutations. Evidence source is OTHER
      because this is a biochemical mechanism paper.
pathophysiology:
- name: MYH7 Missense Variant in Beta-Cardiac Myosin Heavy Chain
  biological_scale: MOLECULAR
  role: trigger
  conforms_to: "cardiomyopathy_maladaptive_remodeling#Primary Cardiomyocyte Insult"
  description: >-
    The initiating lesion in CMH1 is a heterozygous missense variant in MYH7,
    the gene encoding the beta-cardiac myosin heavy chain — the ATP-hydrolysing
    motor of the sarcomeric thick filament. The founding example, and still the
    archetype, is Arg403Gln (R403Q), the first mutation ever linked to familial
    hypertrophic cardiomyopathy; R403Q sits in the globular myosin head (S1),
    the region responsible for actin interaction and motor function. In contrast
    to the truncating, haploinsufficiency mechanism of MYBPC3 disease, most MYH7
    alleles are missense changes that are stably expressed and incorporated into
    the sarcomere as a mutant motor, so the mutant protein perturbs contraction
    directly rather than by its absence.
  genes:
  - preferred_term: MYH7
    term:
      id: hgnc:7577
      label: MYH7
  cell_types:
  - preferred_term: Cardiomyocyte
    term:
      id: CL:0000746
      label: cardiac muscle cell
  biological_processes:
  - preferred_term: Sarcomere Organization
    term:
      id: GO:0045214
      label: sarcomere organization
    modifier: ABNORMAL
  cellular_components:
  - preferred_term: Myosin thick filament
    term:
      id: GO:0032982
      label: myosin filament
  evidence:
  - reference: PMID:1975517
    reference_title: "A molecular basis for familial hypertrophic cardiomyopathy: a beta cardiac myosin heavy chain gene missense mutation."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      A point mutation in exon 13 of the beta cardiac myosin heavy chain (MHC)
      gene is present in all individuals affected with familial hypertrophic
      cardiomyopathy (FHC) from a large kindred. This missense mutation converts
      a highly conserved arginine residue (Arg-403) to a glutamine.
    explanation: >-
      The founding CMH1 report: co-segregation of the MYH7 R403Q missense variant
      with familial hypertrophic cardiomyopathy in a large kindred, establishing
      the beta-cardiac myosin heavy chain gene as a cause of the disease.
  - reference: PMID:23798412
    reference_title: "Molecular consequences of the R453C hypertrophic cardiomyopathy mutation on human β-cardiac myosin motor function."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      HCM is caused by mutations in the genes encoding the fundamental
      force-generating machinery of the cardiac muscle, including β-cardiac
      myosin.
    explanation: >-
      Places the MYH7 lesion in the force-generating machinery of the sarcomere,
      the trigger for this entity.
  downstream:
  - target: Destabilization of the Autoinhibited Super-Relaxed State
    causal_link_type: DIRECT
    description: >-
      The mutant motor, incorporated into the thick filament, destabilizes the
      folded-back autoinhibited state that normally sequesters myosin heads.
- name: Destabilization of the Autoinhibited Super-Relaxed State
  biological_scale: MOLECULAR
  role: amplifier
  description: >-
    In the resting sarcomere a large fraction of myosin heads occupy the
    super-relaxed (SRX) state — folded back against the thick-filament backbone
    in the interacting-heads motif (IHM), with very low ATPase activity, forming
    an energy-conserving reserve of motors withheld from force generation. Many
    MYH7 HCM mutations map to the interfaces that stabilize this folded-back
    state (the myosin mesa, the head-head and head-tail contacts), and they
    weaken those intramolecular interactions. The result is release of
    sequestered heads from the SRX/IHM into the disordered-relaxed state, so more
    heads become functionally accessible to actin — the molecular origin of
    hypercontractility shared across a large subset of MYH7 alleles.
  cell_types:
  - preferred_term: Cardiomyocyte
    term:
      id: CL:0000746
      label: cardiac muscle cell
  molecular_functions:
  - preferred_term: ATP Hydrolysis by Myosin
    term:
      id: GO:0016887
      label: ATP hydrolysis activity
    modifier: INCREASED
  cellular_components:
  - preferred_term: Myosin thick filament
    term:
      id: GO:0032982
      label: myosin filament
  evidence:
  - reference: PMID:31213605
    reference_title: "β-Cardiac myosin hypertrophic cardiomyopathy mutations release sequestered heads and increase enzymatic activity."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      folded-back sequestered state referred to as the interacting heads motif
      (IHM) lead to a significant increase in the number of heads functionally
      accessible for interaction with actin.
    explanation: >-
      Direct biochemical evidence that MYH7 HCM mutations at IHM interfaces
      release sequestered heads, increasing the number available to interact with
      actin.
  - reference: PMID:28481356
    reference_title: "The myosin mesa and the basis of hypercontractility caused by hypertrophic cardiomyopathy mutations."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      hypercontractility is due to an increase in the number of myosin heads (S1)
      that are accessible for force production.
    explanation: >-
      States the core hypothesis this node captures: HCM hypercontractility
      arises from an increase in the number of force-producing heads freed from
      the sequestered state.
  - reference: PMID:34117120
    reference_title: "Hypertrophic cardiomyopathy β-cardiac myosin mutation (P710R) leads to hypercontractility by disrupting super relaxed state."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      destabilized the super relaxed state in longer, two-headed myosin
      constructs, freeing more heads to generate force.
    explanation: >-
      Shows a specific MYH7 mutation (P710R) destabilizing the super-relaxed
      state and freeing heads for force generation.
  - reference: PMID:40118457
    reference_title: "Dynamics of β-cardiac myosin between the super-relaxed and disordered-relaxed states."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      Some mutations which cause hypertrophic or dilated cardiomyopathies alter
      the SRX-DRX equilibrium, but not all mutations.
    explanation: >-
      Important caveat curated with the node: SRX-DRX destabilization is a common
      but not universal consequence of cardiomyopathy myosin mutations, so this
      arm applies to the large subset of alleles that shift the equilibrium
      rather than to every MYH7 variant. PARTIAL for that reason.
  downstream:
  - target: Increased Motor Force and Energetic Cost
    causal_link_type: DIRECT
    description: >-
      Heads released from the super-relaxed reserve, together with allele-specific
      changes in the motor stroke, raise ensemble force and the ATP cost of
      contraction.
- name: Increased Motor Force and Energetic Cost
  biological_scale: MOLECULAR
  role: effector
  description: >-
    Beyond releasing sequestered heads, individual MYH7 missense alleles alter
    the mechanochemistry of the motor stroke itself. Single-molecule and
    myofibril studies show mutation-specific changes — increased intrinsic force
    per cross-bridge, and faster cross-bridge detachment kinetics — that together
    raise the force generated by the ensemble of motors and increase the ATP
    consumed per unit tension (tension cost). The energetic penalty, measured
    directly in human myocardium carrying R403Q, is a distinctive feature of the
    beta-myosin motor lesion and links the molecular defect to the myocardial
    energy deficit seen in HCM.
  cell_types:
  - preferred_term: Cardiomyocyte
    term:
      id: CL:0000746
      label: cardiac muscle cell
  molecular_functions:
  - preferred_term: ATP Hydrolysis by Myosin
    term:
      id: GO:0016887
      label: ATP hydrolysis activity
    modifier: INCREASED
  biological_processes:
  - preferred_term: Actin-Myosin Filament Sliding
    term:
      id: GO:0033275
      label: actin-myosin filament sliding
    modifier: ABNORMAL
  evidence:
  - reference: PMID:23798412
    reference_title: "Molecular consequences of the R453C hypertrophic cardiomyopathy mutation on human β-cardiac myosin motor function."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      The major change in the R453C human β-cardiac subfragment 1 is a 50%
      increase in the intrinsic force of the motor compared with wild type, with
      no appreciable change in the stroke size, as observed with a dual-beam
      optical trap.
    explanation: >-
      Single-molecule optical-trap measurement of a raised intrinsic force for an
      MYH7 mutant motor, predicting a hypercontractile ensemble.
  - reference: PMID:23798412
    reference_title: "Molecular consequences of the R453C hypertrophic cardiomyopathy mutation on human β-cardiac myosin motor function."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      Overall, this study suggests that the R453C mutation should result in a
      hypercontractile state in the heart muscle.
    explanation: >-
      States the functional conclusion — a hypercontractile state — that the
      motor-level force change produces.
  - reference: PMID:24928957
    reference_title: "Faster cross-bridge detachment and increased tension cost in human hypertrophic cardiomyopathy with the R403Q MYH7 mutation."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      This correlation suggests that faster cross-bridge relaxation kinetics
      results in an increase in energetic cost of tension generation in human HCM
      with the R403Q mutation compared to HCMsmn.
    explanation: >-
      Direct measurement in human R403Q myocardium (single myofibrils and muscle
      strips) linking faster cross-bridge kinetics to an increased energetic cost
      of tension generation, evidence obtained in patient cardiac tissue.
  downstream:
  - target: Sarcomere Hypercontractility with Impaired Relaxation
    causal_link_type: DIRECT
    description: >-
      Raised ensemble force and disturbed cross-bridge kinetics translate into
      cellular hypercontractility with impaired diastolic relaxation.
- name: Sarcomere Hypercontractility with Impaired Relaxation
  biological_scale: CELLULAR
  role: effector
  description: >-
    At the cell level the molecular changes converge on hypercontractility:
    CRISPR-edited human iPSC-derived cardiomyocytes carrying an MYH7 HCM mutation
    generate significantly greater contractile force than isogenic controls.
    Because the same lesion disturbs cross-bridge relaxation kinetics, systolic
    hypercontraction is accompanied by impaired diastolic relaxation. This
    hypercontractile, poorly relaxing cell state — a key pathophysiological
    abnormality of HCM and the determinant of dynamic outflow obstruction — is
    the cellular phenotype the cardiac myosin inhibitors were designed to
    normalize.
  cell_types:
  - preferred_term: Ventricular cardiomyocyte
    term:
      id: CL:0002131
      label: regular ventricular cardiac myocyte
  biological_processes:
  - preferred_term: Cardiac Muscle Cell Contraction
    term:
      id: GO:0086003
      label: cardiac muscle cell contraction
    modifier: INCREASED
  - preferred_term: Regulation of Cardiac Muscle Contraction
    term:
      id: GO:0055117
      label: regulation of cardiac muscle contraction
    modifier: ABNORMAL
  evidence:
  - reference: PMID:34117120
    reference_title: "Hypertrophic cardiomyopathy β-cardiac myosin mutation (P710R) leads to hypercontractility by disrupting super relaxed state."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      produced significantly increased force (measured by traction force
      microscopy) compared with isogenic control cells.
    explanation: >-
      Isogenic human iPSC-cardiomyocyte evidence that an MYH7 HCM mutation
      increases cellular contractile force.
  - reference: PMID:32871100
    reference_title: "Mavacamten for treatment of symptomatic obstructive hypertrophic cardiomyopathy (EXPLORER-HCM): a randomised, double-blind, placebo-controlled, phase 3 trial."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Cardiac muscle hypercontractility is a key pathophysiological abnormality
      in hypertrophic cardiomyopathy, and a major determinant of dynamic left
      ventricular outflow tract (LVOT) obstruction.
    explanation: >-
      Confirms hypercontractility as the central pathophysiological abnormality
      of HCM and links it to outflow obstruction; PARTIAL because the trial
      enrolled obstructive HCM broadly rather than genotyped MYH7 carriers.
  downstream:
  - target: Cell-to-Cell Contractile Imbalance
    causal_link_type: DIRECT
    description: >-
      Because mutant and wild-type MYH7 alleles are transcribed in variable
      proportions between neighboring myocytes, hypercontractility is unevenly
      distributed across the myocardium.
  - target: Ventricular Hypertrophy, Myocyte Disarray and Fibrosis
    causal_link_type: DIRECT
    description: >-
      Sustained hypercontractility with raised energetic cost drives the
      structural remodeling of the ventricle.
- name: Cell-to-Cell Contractile Imbalance
  biological_scale: CELLULAR
  role: amplifier
  description: >-
    A mechanism specific to the missense/poison-peptide biology of MYH7 disease.
    Because the mutant and wild-type MYH7 alleles are transcribed in a random,
    burst-like fashion and independently of one another, the fraction of mutant
    beta-myosin varies markedly from cardiomyocyte to cardiomyocyte within a
    single patient's myocardium. This produces much greater cell-to-cell
    variability of contractile function than in control hearts. The resulting
    mechanical mismatch between adjacent, differently loaded myocytes is proposed
    to drive the myofiber disarray and interstitial fibrosis that are hallmarks
    of HCM, providing a route from the molecular lesion to the tissue phenotype
    that does not depend on the average level of hypercontractility alone.
  cell_types:
  - preferred_term: Cardiomyocyte
    term:
      id: CL:0000746
      label: cardiac muscle cell
  biological_processes:
  - preferred_term: Cardiac Muscle Cell Contraction
    term:
      id: GO:0086003
      label: cardiac muscle cell contraction
    modifier: ABNORMAL
  evidence:
  - reference: PMID:30740621
    reference_title: "Altered force generation and cell-to-cell contractile imbalance in hypertrophic cardiomyopathy."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      substantial contractile variability from cardiomyocyte to cardiomyocyte
      within a patient's myocardium, much higher than in controls.
    explanation: >-
      Documents the cell-to-cell contractile variability in beta-MyHC-mutant
      patient myocardium that this node captures.
  - reference: PMID:30740621
    reference_title: "Altered force generation and cell-to-cell contractile imbalance in hypertrophic cardiomyopathy."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      contractile imbalance due to unequal fractions of mutated and wildtype
      protein among individual cardiomyocytes over time will induce cardiomyocyte
      disarray and fibrosis, hallmarks of HCM.
    explanation: >-
      States the contractile-imbalance hypothesis linking unequal mutant-allele
      expression to disarray and fibrosis, the hallmarks of HCM.
  downstream:
  - target: Ventricular Hypertrophy, Myocyte Disarray and Fibrosis
    causal_link_type: INDIRECT_UNKNOWN_INTERMEDIATES
    description: >-
      Mechanical mismatch between neighboring myocytes promotes disarray and
      fibrotic remodeling.
- name: Ventricular Hypertrophy, Myocyte Disarray and Fibrosis
  biological_scale: TISSUE
  role: central_effector
  conforms_to: "cardiomyopathy_maladaptive_remodeling#Ventricular Remodeling"
  description: >-
    Chronic hypercontractility with raised energetic cost, amplified by
    cell-to-cell contractile imbalance, drives the structural remodeling that
    defines the clinical phenotype: cardiomyocyte hypertrophy — typically
    asymmetric and septal-predominant — together with myofiber disarray and
    interstitial and replacement fibrosis. Pro-hypertrophic growth signalling
    contributes: in an MYH7-mutant human iPSC-cardiomyocyte model, cellular
    hypertrophy was prevented by inhibition of ERK or Akt, implicating those
    pathways downstream of the mechanical stimulus. Fibrosis provides the
    substrate for both diastolic dysfunction and re-entrant arrhythmia.
  cell_types:
  - preferred_term: Cardiomyocyte
    term:
      id: CL:0000746
      label: cardiac muscle cell
  - preferred_term: Cardiac Fibroblast
    term:
      id: CL:0002548
      label: fibroblast of cardiac tissue
  biological_processes:
  - preferred_term: Cardiac Muscle Hypertrophy in Response to Stress
    term:
      id: GO:0014898
      label: cardiac muscle hypertrophy in response to stress
    modifier: INCREASED
  - preferred_term: Extracellular Matrix Organization
    term:
      id: GO:0030198
      label: extracellular matrix organization
    modifier: INCREASED
  locations:
  - preferred_term: Interventricular septum
    term:
      id: UBERON:0002094
      label: interventricular septum
  - preferred_term: Left ventricle
    term:
      id: UBERON:0002084
      label: heart left ventricle
  evidence:
  - reference: PMID:28912181
    reference_title: "Hypertrophic Cardiomyopathy: Genetics, Pathogenesis, Clinical Manifestations, Diagnosis, and Therapy."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: "The histological features of HCM include myocyte hypertrophy and disarray, as well as interstitial fibrosis."
    explanation: >-
      Names the defining tissue-level triad — myocyte hypertrophy, disarray, and
      interstitial fibrosis — that this node represents.
  - reference: PMID:34117120
    reference_title: "Hypertrophic cardiomyopathy β-cardiac myosin mutation (P710R) leads to hypercontractility by disrupting super relaxed state."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: "Cellular hypertrophy was prevented in the P710R cells by inhibition of ERK or Akt."
    explanation: >-
      Implicates ERK and Akt growth signalling downstream of the MYH7 mechanical
      lesion in producing the cellular hypertrophy of this node.
  - reference: PMID:30740621
    reference_title: "Altered force generation and cell-to-cell contractile imbalance in hypertrophic cardiomyopathy."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      contractile imbalance due to unequal fractions of mutated and wildtype
      protein among individual cardiomyocytes over time will induce cardiomyocyte
      disarray and fibrosis, hallmarks of HCM.
    explanation: >-
      Ties the upstream contractile imbalance to the disarray and fibrosis of
      this tissue node.
  downstream:
  - target: Diastolic Dysfunction, Outflow Obstruction, Heart Failure and Arrhythmic Risk
    causal_link_type: DIRECT
- name: Diastolic Dysfunction, Outflow Obstruction, Heart Failure and Arrhythmic Risk
  biological_scale: ORGANISM
  role: consequence
  conforms_to: "cardiomyopathy_maladaptive_remodeling#Structural Cardiac Impairment and Heart Failure"
  description: >-
    The hypertrophied, disarrayed and fibrotic ventricle produces the clinical
    endpoints of CMH1. Hypercontractility with septal hypertrophy generates
    dynamic left ventricular outflow tract obstruction in a subset of patients;
    the stiff, poorly relaxing ventricle causes diastolic dysfunction; and the
    fibrotic, electrically heterogeneous substrate carries a risk of malignant
    ventricular arrhythmia and sudden cardiac death that is clinically important
    for MYH7 variants. Sarcomere-positive disease, of which MYH7 is a leading
    cause, presents younger and with a stronger family history of sudden death
    than sarcomere-negative HCM.
  cell_types:
  - preferred_term: Cardiomyocyte
    term:
      id: CL:0000746
      label: cardiac muscle cell
  biological_processes:
  - preferred_term: Heart Contraction
    term:
      id: GO:0060047
      label: heart contraction
    modifier: ABNORMAL
  evidence:
  - reference: PMID:23674365
    reference_title: "A systematic review and meta-analysis of genotype-phenotype associations in patients with hypertrophic cardiomyopathy caused by sarcomeric protein mutations."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      There were no differences when the two most frequently affected genes,
      MYBPC3 and MYH7, were compared.
    explanation: >-
      Records the important nuance that, at the cohort level, MYH7 and MYBPC3 do
      not differ significantly across the pooled clinical features, so MYH7 is
      not simply a uniformly more severe genotype than MYBPC3; it is one of the
      two commonest sarcomeric causes.
  - reference: PMID:39681440
    reference_title: "[Correlation between genotype and clinical phenotype in hypertrophic cardiomyopathy families with MYH7-R453C mutation]."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      In the five families with HCM carrying MYH7-R453C mutations, genotype is
      highly correlated with clinical phenotype, and patients have a high risk of
      sudden death and poor prognosis.
    explanation: >-
      A family study of a specific MYH7 variant documenting high sudden-death
      risk and poor prognosis, the arrhythmic endpoint of this node.
phenotypes:
- name: Hypertrophic Cardiomyopathy
  category: Cardiovascular
  description: >-
    Unexplained left ventricular hypertrophy, not accounted for by abnormal
    loading conditions, is the defining feature of CMH1.
  phenotype_term:
    preferred_term: Hypertrophic cardiomyopathy
    term:
      id: HP:0001639
      label: Hypertrophic cardiomyopathy
  frequency: OBLIGATE
  evidence:
  - reference: PMID:28912181
    reference_title: "Hypertrophic Cardiomyopathy: Genetics, Pathogenesis, Clinical Manifestations, Diagnosis, and Therapy."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: "Hypertrophic cardiomyopathy (HCM) is a genetic disorder that is characterized by left ventricular hypertrophy unexplained by secondary causes and a nondilated left ventricle with preserved or increased ejection fraction."
    explanation: >-
      States the defining clinical feature — unexplained left ventricular
      hypertrophy with a nondilated ventricle — of the disease this MYH7 form
      manifests.
- name: Asymmetric Septal Hypertrophy
  category: Cardiovascular
  description: >-
    Asymmetric, septal-predominant left ventricular hypertrophy is the classic
    morphological pattern of MYH7-related hypertrophic cardiomyopathy.
  phenotype_term:
    preferred_term: Asymmetric septal hypertrophy
    term:
      id: HP:0001670
      label: Asymmetric septal hypertrophy
  evidence:
  - reference: PMID:23674365
    reference_title: "A systematic review and meta-analysis of genotype-phenotype associations in patients with hypertrophic cardiomyopathy caused by sarcomeric protein mutations."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      The presence of any sarcomere gene mutation was associated with a younger
      age at presentation (38.4 vs 46.0 years, p<0.0005), a family history of HCM
      (50.6% vs 23.1%, p<0.0005), a family history of SCD (27.0% vs 14.9%,
      p<0.0005) and greater MLVWT (21.0 vs 19.3 mm, p=0.03).
    explanation: >-
      Sarcomere-positive HCM, of which MYH7 is a leading cause, shows greater
      maximum left ventricular wall thickness than sarcomere-negative disease,
      supporting the pronounced hypertrophy of this member.
- name: Left Ventricular Outflow Tract Obstruction
  category: Cardiovascular
  description: >-
    Dynamic obstruction of the left ventricular outflow tract arises from
    hypercontractility combined with septal hypertrophy and systolic anterior
    motion of the mitral valve, defining the obstructive phenotype targeted by
    myosin inhibitors and septal reduction therapy.
  phenotype_term:
    preferred_term: Left ventricular outflow tract obstruction
    term:
      id: HP:0032092
      label: Left ventricular outflow tract obstruction
  evidence:
  - reference: PMID:32871100
    reference_title: "Mavacamten for treatment of symptomatic obstructive hypertrophic cardiomyopathy (EXPLORER-HCM): a randomised, double-blind, placebo-controlled, phase 3 trial."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Cardiac muscle hypercontractility is a key pathophysiological abnormality
      in hypertrophic cardiomyopathy, and a major determinant of dynamic left
      ventricular outflow tract (LVOT) obstruction.
    explanation: >-
      Links hypercontractility to dynamic LVOT obstruction; PARTIAL because the
      trial population is obstructive HCM broadly rather than genotyped MYH7
      carriers.
- name: Left Ventricular Diastolic Dysfunction
  category: Cardiovascular
  description: >-
    Impaired relaxation and increased chamber stiffness produce diastolic
    dysfunction, an early and characteristic functional consequence of the
    hypercontractile, poorly relaxing myocardium.
  phenotype_term:
    preferred_term: Left ventricular diastolic dysfunction
    term:
      id: HP:0025168
      label: Left ventricular diastolic dysfunction
  evidence:
  - reference: PMID:24928957
    reference_title: "Faster cross-bridge detachment and increased tension cost in human hypertrophic cardiomyopathy with the R403Q MYH7 mutation."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Cross-bridge slow relaxation kinetics in single R403Q myofibrils was
      significantly higher (P < 0.0001) than in HCMsmn myofibrils
    explanation: >-
      Documents disturbed cross-bridge relaxation kinetics in human R403Q
      myocardium, the molecular basis of the impaired relaxation underlying
      diastolic dysfunction.
- name: Sudden Cardiac Death
  category: Cardiovascular
  description: >-
    Malignant ventricular arrhythmia on the fibrotic, hypertrophied substrate
    can cause sudden cardiac death, sometimes as the presenting event; the risk
    is clinically important for MYH7 variants and family history of sudden death
    is characteristic.
  phenotype_term:
    preferred_term: Sudden cardiac death
    term:
      id: HP:0001645
      label: Sudden cardiac death
  evidence:
  - reference: PMID:39681440
    reference_title: "[Correlation between genotype and clinical phenotype in hypertrophic cardiomyopathy families with MYH7-R453C mutation]."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      In the five families with HCM carrying MYH7-R453C mutations, genotype is
      highly correlated with clinical phenotype, and patients have a high risk of
      sudden death and poor prognosis.
    explanation: >-
      A family study of an MYH7 variant documenting a high risk of sudden death.
- name: Ventricular Arrhythmia
  category: Cardiovascular
  description: >-
    The electrically heterogeneous, fibrotic myocardium supports malignant
    ventricular arrhythmia, the proximate cause of sudden cardiac death in this
    disease.
  phenotype_term:
    preferred_term: Ventricular arrhythmia
    term:
      id: HP:0004308
      label: Ventricular arrhythmia
  evidence:
  - reference: PMID:39681440
    reference_title: "[Correlation between genotype and clinical phenotype in hypertrophic cardiomyopathy families with MYH7-R453C mutation]."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      2 experienced (F1Ⅲ1, F3Ⅲ3) events of sudden cardiac death survival
    explanation: >-
      Aborted sudden cardiac death (survived cardiac arrest) in MYH7-R453C
      carriers evidences the malignant ventricular arrhythmia of this node.
- name: Congestive Heart Failure
  category: Cardiovascular
  description: >-
    Progressive diastolic dysfunction, and in a minority progression toward
    systolic impairment or an end-stage phase, produces heart failure symptoms.
  phenotype_term:
    preferred_term: Congestive heart failure
    term:
      id: HP:0001635
      label: Congestive heart failure
  evidence:
  - reference: PMID:39681440
    reference_title: "[Correlation between genotype and clinical phenotype in hypertrophic cardiomyopathy families with MYH7-R453C mutation]."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      2(F1Ⅱ2, F3Ⅱ1) died from heart failure during the follow-up period.
    explanation: >-
      Deaths from heart failure among MYH7-R453C carriers evidence heart failure
      as a clinical endpoint of the disease.
genetic:
- name: MYH7
  gene_term:
    preferred_term: MYH7
    term:
      id: hgnc:7577
      label: MYH7
  relationship_type: CAUSATIVE
  frequency: >-
    With MYBPC3, one of the two commonest genetic causes of hypertrophic
    cardiomyopathy: nearly 40% of identified HCM-causing mutations fall in
    beta-cardiac myosin.
  case_fractions:
  - population: Identified HCM-causing mutations (genotyped probands)
    case_fraction_percent: 40.0
    notes: >-
      This is a share of identified HCM-causing mutations, not a share of all
      hypertrophic cardiomyopathy cases. The two differ because a substantial
      fraction of clinically diagnosed HCM is genotype-negative, so the figure
      overstates the share of the whole disease population and is recorded
      with that denominator named explicitly.
    evidence:
    - reference: PMID:31213605
      reference_title: "β-Cardiac myosin hypertrophic cardiomyopathy mutations release sequestered heads and increase enzymatic activity."
      supports: SUPPORT
      evidence_source: OTHER
      snippet: >-
        Nearly 40 percent of HCM-causing mutations are found in human
        β-cardiac myosin.
      explanation: >-
        Directly states the beta-cardiac myosin share of HCM-causing
        mutations.
  evidence:
  - reference: CGGV:assertion_31325c90-05cb-4db0-9372-e4f705cd5c82-2023-07-12T160000.000Z
    reference_title: "MYH7 / hypertrophic cardiomyopathy (Definitive)"
    supports: SUPPORT
    evidence_source: OTHER
    snippet: >-
      MYH7 | HGNC:7577 | hypertrophic cardiomyopathy | MONDO:0005045 | AD |
      Definitive
    explanation: >-
      ClinGen's Gene Curation Expert Panel classifies this gene-disease
      relationship as Definitive, which is the authority for curating the gene
      as CAUSATIVE rather than a candidate.
  - reference: PMID:23674365
    reference_title: "A systematic review and meta-analysis of genotype-phenotype associations in patients with hypertrophic cardiomyopathy caused by sarcomeric protein mutations."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      The presence of any sarcomere gene mutation was associated with a
      younger age at presentation (38.4 vs 46.0 years, p<0.0005), a family
      history of HCM (50.6% vs 23.1%, p<0.0005), a family history of SCD
      (27.0% vs 14.9%, p<0.0005) and greater MLVWT (21.0 vs 19.3 mm, p=0.03).
    explanation: >-
      The cohort is sarcomere-mutation carriers across genes rather than MYH7
      carriers, so this is class-level evidence that a sarcomere genotype
      shifts age at presentation and family history. Curated PARTIAL for that
      reason; the same study's finding of no outcome difference between MYH7
      and MYBPC3 carriers is recorded in this entry's pathophysiology.
treatments:
- name: Mavacamten (Cardiac Myosin Inhibitor)
  description: >-
    Mavacamten is a small-molecule allosteric inhibitor of cardiac myosin ATPase
    that reduces the number of force-generating actin-myosin cross-bridges,
    stabilizing myosin in the energy-sparing super-relaxed state. It directly
    counteracts the hypercontractility that is the central abnormality of HCM, and
    is approved for symptomatic obstructive hypertrophic cardiomyopathy; because
    beta-cardiac myosin is the MYH7 gene product, the drug acts on the same motor
    that carries the CMH1 lesion.
  therapeutic_modality: SMALL_MOLECULE
  treatment_term:
    preferred_term: Pharmacotherapy
    term:
      id: NCIT:C15986
      label: Pharmacotherapy
    therapeutic_agent:
    - preferred_term: mavacamten
      term:
        id: CHEBI:756998
        label: mavacamten
  target_mechanisms:
  - target: Destabilization of the Autoinhibited Super-Relaxed State
    treatment_effect: INHIBITS
    description: >-
      Mavacamten stabilizes the super-relaxed state and reduces the number of
      heads available for force generation, directly opposing the SRX
      destabilization that drives hypercontractility.
    evidence:
    - reference: PMID:40118457
      reference_title: "Dynamics of β-cardiac myosin between the super-relaxed and disordered-relaxed states."
      supports: SUPPORT
      evidence_source: IN_VITRO
      snippet: >-
        cardiac myosin inhibitor mavacamten slows nucleotide release by an equal
        factor for both heavy meromyosin and subfragment 1, thus only indirectly
        influencing the occupancy time of the SRX state.
      explanation: >-
        Establishes mavacamten's biochemical action on beta-cardiac myosin
        nucleotide handling and its effect on SRX-state occupancy, the node it
        targets.
  evidence:
  - reference: PMID:32871100
    reference_title: "Mavacamten for treatment of symptomatic obstructive hypertrophic cardiomyopathy (EXPLORER-HCM): a randomised, double-blind, placebo-controlled, phase 3 trial."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Cardiac muscle hypercontractility is a key pathophysiological abnormality
      in hypertrophic cardiomyopathy, and a major determinant of dynamic left
      ventricular outflow tract (LVOT) obstruction.
    explanation: >-
      The pivotal phase 3 trial establishing the rationale and use of a cardiac
      myosin inhibitor targeting hypercontractility in obstructive HCM.
- name: Septal Reduction (Surgical Myectomy)
  description: >-
    Surgical septal myectomy relieves severe drug-refractory left ventricular
    outflow tract obstruction by resecting hypertrophied basal septal muscle.
  therapeutic_modality: SURGERY
  treatment_term:
    preferred_term: septal myectomy
    term:
      id: NCIT:C51591
      label: Myectomy
- name: Implantable Cardioverter-Defibrillator
  description: >-
    An implantable cardioverter-defibrillator provides secondary or primary
    prevention of sudden cardiac death in patients judged to be at high arrhythmic
    risk, terminating malignant ventricular arrhythmia.
  therapeutic_modality: DEVICE
  treatment_term:
    preferred_term: implantable cardioverter-defibrillator placement
    term:
      id: NCIT:C80435
      label: Implantable Cardioverter-Defibrillator Placement
clinical_trials:
- name: NCT01912534
  phase: PHASE_II
  status: COMPLETED
  description: >-
    VANISH (Valsartan for Attenuating Disease Evolution in Early Sarcomeric
    Hypertrophic Cardiomyopathy) tested whether the angiotensin receptor blocker
    valsartan can modify disease progression in young carriers of sarcomeric gene
    mutations, including MYH7, with early-stage HCM.
  target_phenotypes:
  - preferred_term: Hypertrophic cardiomyopathy
    term:
      id: HP:0001639
      label: Hypertrophic cardiomyopathy
  evidence:
  - reference: PMID:31813281
    reference_title: "Baseline Characteristics of the VANISH Cohort."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      targeted young sarcomeric gene mutation carriers with early-stage
      hypertrophic cardiomyopathy (HCM) to test whether valsartan can modify
      disease progression.
    explanation: >-
      Describes the VANISH trial's design and its enrollment of young sarcomeric
      (including MYH7) mutation carriers with early HCM.
references:
- reference: PMID:20301725
  title: Nonsyndromic Hypertrophic Cardiomyopathy Overview.
  tags:
  - GeneReviews
notes: >-
  Scope and mechanism contrast with CMH4 (MYBPC3). CMH1 is curated as the
  MYH7/beta-cardiac myosin form because MYH7 is, with MYBPC3, one of the two
  commonest genetic causes of HCM and was the founding molecular diagnosis of the
  field (R403Q, 1990). It is deliberately contrasted with the MYBPC3 entry: MYBPC3
  disease is predominantly truncating and acts by haploinsufficiency (loss of the
  cMyBP-C brake), whereas MYH7 disease is predominantly missense and acts by a
  gain of contractile function in a stably expressed mutant motor (super-relaxed
  state destabilization, raised intrinsic force, increased tension cost). Both
  converge on the same maladaptive-remodeling module.

  Allele-heterogeneity caveat preserved on the pathograph: not every MYH7 mutation
  shifts the SRX-DRX equilibrium (PMID:40118457), so the super-relaxed-state arm
  is curated as the mechanism of a large subset of alleles rather than a universal
  one, and the motor-force arm (intrinsic force, tension cost) is curated
  separately. At the cohort level MYH7 and MYBPC3 do not differ significantly
  across pooled clinical features (PMID:23674365), so the entry does not assert
  MYH7 as a uniformly more severe genotype.

  Curating this entry deletes the stub stubs/Hypertrophic_Cardiomyopathy_1.yaml
  and closes the highest-value gap named in the Familial Hypertrophic
  Cardiomyopathy grouping notes. This entry is added as a member of that grouping.

  GeneReviews scope. The GeneReviews resource applicable to this entry is the
  disease-level "Nonsyndromic Hypertrophic Cardiomyopathy Overview"
  (PMID:20301725), tagged accordingly in `references`. Its indexed PubMed record
  is content_type abstract_only and carries only the chapter's purpose
  statement, not the Clinical Characteristics, Management, or Genetic Counseling
  sections, so section-by-section GeneReviews mining is not possible from the
  cache and no snippet is quoted from it. The clinical-characteristics baseline
  for this entry is therefore built from the primary MYH7 cohort and pedigree
  literature cited throughout, principally PMID:28912181, PMID:23674365,
  PMID:32871100, PMID:24928957, PMID:39681440.
📚

References & Deep Research

References

1
Nonsyndromic Hypertrophic Cardiomyopathy Overview.
No top-level findings curated for this source.