Congenital Sodium Diarrhea

Congenital sodium diarrhea (CSD) is a rare congenital diarrhea and enteropathy (CODE) defined by perinatal-onset watery diarrhea with disproportionately high fecal sodium loss, hyponatremia and metabolic ACIDOSIS. With congenital chloride diarrhea it forms the electrolyte-transport-related class of the CODEs - disorders in which the failed apical transport step handles an ion rather than an ingested nutrient, so the diarrhea is independent of feeding and cannot be treated by eliminating any dietary constituent. CSD is genetically heterogeneous but mechanistically convergent. Classic (non-syndromic) CSD is caused by recessive loss-of-function variants in SLC9A3, encoding NHE3, the major sodium/proton exchanger of the intestinal brush border. A second non-syndromic form is caused by dominant ACTIVATING variants in GUCY2C, encoding guanylate cyclase C: excess cyclic GMP both drives CFTR-mediated chloride and water secretion and inhibits NHE3, which is itself a downstream target of GC-C - so the gain-of-function and loss-of-function routes converge on the same failure of NHE3-mediated sodium absorption. A syndromic form is caused by recessive loss-of-function variants in SPINT2, encoding a Kunitz-type serine protease inhibitor, and is distinguished by choanal or anal atresia, hypertelorism and corneal erosions. CSD is the acid-base mirror image of congenital chloride diarrhea: losing the sodium/proton exchanger means protons are retained rather than secreted, giving acidosis where loss of the chloride/bicarbonate exchanger gives alkalosis. Mortality is high in the first years from electrolyte imbalance and the complications of parenteral nutrition, but survivors can adapt to partial or complete enteral nutrition - a natural history distinctly better than that of the structural enteropathies.

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2
Inheritance
6
Pathophys.
7
Phenotypes
7
Pathograph
3
Genes
1
Medical Actions
3
Subtypes
šŸ‘Ŗ

Inheritance

2
Autosomal recessive inheritance HP:0000007
Classic (SLC9A3) and syndromic (SPINT2) CSD are autosomal recessive. Consanguinity is common in reported kindreds; the original Austrian cohort traced five affected infants in a circumscribed rural area to a single common ancestor five generations back.
Autosomal recessive inheritance
Show evidence (2 references)
PMID:11113072 SUPPORT Human Clinical
"Pedigree analysis of the affected 2 CSD families revealed parental consanguinity and a common single ancestor 5 generations ago."
Pedigree evidence for recessive inheritance with a founder effect in a geographically circumscribed population.
PMID:19185281 SUPPORT Human Clinical
"Autosomal-recessive congenital sodium diarrhea (CSD) is characterized by perinatal onset of a persistent watery diarrhea with nonproportionally high fecal sodium excretion."
States the recessive inheritance and the defining clinical and biochemical features of the disorder.
Autosomal dominant inheritance HP:0000006
The GUCY2C-associated form is autosomal dominant and results from activating (gain-of-function) rather than loss-of-function variants, so it does not follow the recessive pattern of the other two forms. It was delineated in a 32-member Norwegian family. Applies to the GUCY2C-activating subtype only.
Autosomal dominant inheritance
Show evidence (1 reference)
PMID:22436048 SUPPORT Human Clinical
"Familial diarrhea disorders are, in most cases, severe and caused by recessive mutations. We describe the cause of a novel dominant disease in 32 members of a Norwegian family."
Establishes the dominant inheritance of the GUCY2C form and explicitly contrasts it with the recessive majority of familial diarrheas.
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Subtypes

3
Classic (non-syndromic) congenital sodium diarrhea, SLC9A3-related
Recessive SLC9A3/NHE3 loss of function. Isolated intestinal disease with no atresias or facial features. Presented in seven families in the cohort that delineated the syndromic/classic split.
Syndromic congenital sodium diarrhea, SPINT2-related
Recessive SPINT2 loss of function. Intestinal disease plus choanal or anal atresia, hypertelorism and corneal erosions. Occurred in ten families in the delineating cohort. SPINT2 also causes a syndromic form of congenital tufting enteropathy.
Familial GUCY2C diarrhea syndrome (dominant, activating)
GUCY2C hgnc:4688 HUGO Gene Nomenclature Committee (hgnc) Relation: this subtype is caused by variation in this gene This subtype is caused by variation in GUCY2C (hgnc:4688). hgnc:4688 is a gene from the HUGO Gene Nomenclature Committee.
Dominant activating GUCY2C variant raising cyclic GMP. Milder, early-onset chronic diarrhea with increased susceptibility to inflammatory bowel disease, small-bowel obstruction and esophagitis. Paradoxically associated with prolonged rather than accelerated gut transit.
Show evidence (1 reference)
PMID:28957388 SUPPORT Human Clinical
"Despite having diarrhea, the FGDS patients have prolonged transit time through the gut compared to HC, particularly in colon."
Supports the paradoxical motility phenotype asserted for this subtype - diarrhea with prolonged rather than accelerated transit, most marked in the colon - which distinguishes it from the other CSD forms.
āš™

Pathophysiology

6
Constitutive Guanylate Cyclase C Activation
The upstream lesion of the GUCY2C arm only. A dominant activating variant in guanylate cyclase C raises cyclic GMP on ligand exposure. Excess cGMP acts in two directions: it hyperactivates CFTR, driving chloride and water secretion, and it inhibits NHE3, which is itself a downstream target of GC-C. This node is therefore placed upstream of the NHE3 lesion rather than beside it - the two are sequential in the GUCY2C arm, not alternative entry points at the same level. Conformers entering through SLC9A3 do not pass through this node.
Small-intestinal enterocyte CL:0000584 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Small-intestinal enterocyte, annotated with enterocyte (CL:0000584). CL:0000584 is a cell type from the Cell Ontology.
Cyclic GMP production GO:0006182 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves increased Cyclic GMP production, annotated with cGMP biosynthetic process (GO:0006182). GO:0006182 is a biological process from the Gene Ontology. ↑ INCREASED
Guanylate cyclase C activity GO:0004383 Gene Ontology (GO) Relation: this pathophysiological event involves this molecular function This pathophysiological event involves Guanylate cyclase C activity, annotated with guanylate cyclase activity (GO:0004383), qualified as gain of function. GO:0004383 is a molecular function from the Gene Ontology. ⇑ GAIN OF FUNCTION
Small intestine UBERON:0002108 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Small intestine (UBERON:0002108). UBERON:0002108 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (2 references)
PMID:22436048 SUPPORT In Vitro
"Exposure of the mutant receptor to its ligands resulted in markedly increased production of cyclic guanosine monophosphate (cGMP)."
Demonstrates the gain-of-function that defines this node - markedly increased cGMP from the mutant receptor. Evidence source is IN_VITRO because cGMP was measured in transfected HEK293T cells.
PMID:22436048 SUPPORT In Vitro
"This may cause hyperactivation of the cystic fibrosis transmembrane regulator (CFTR), leading to increased chloride and water secretion from the enterocytes, and may thus explain the chronic diarrhea in the affected family members."
Gives the parallel secretory arm of this node. The authors hedge it as a proposed explanation, which is why the edge leaving this node is typed as having known intermediates rather than as direct.
Loss of NHE3-Mediated Apical Sodium/Proton Exchange
The convergence point of every genetic form, and the node that carries the transport lesion itself. NHE3 is the major sodium/proton exchanger of the intestinal brush border. It is lost by two distinct routes that meet here rather than being alternatives at the same level: in classic CSD, recessive SLC9A3 variants directly reduce its surface expression or transport function while leaving acute regulation intact; in the GUCY2C form, excess cyclic GMP from the upstream activating variant inhibits it, NHE3 being a downstream target of GC-C. The syndromic SPINT2 form is proposed to act through excess serine protease activity in the affected epithelium, a route that is less well resolved at the transport level and is not modelled as a separate node here.
Small-intestinal enterocyte CL:0000584 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Small-intestinal enterocyte, annotated with enterocyte (CL:0000584). CL:0000584 is a cell type from the Cell Ontology.
Brush-border sodium/proton exchange (NHE3) GO:0015385 Gene Ontology (GO) Relation: this pathophysiological event involves this molecular function This pathophysiological event involves Brush-border sodium/proton exchange (NHE3), annotated with sodium:proton antiporter activity (GO:0015385), qualified as loss of function. GO:0015385 is a molecular function from the Gene Ontology. ⇓ LOSS OF FUNCTION
Enterocyte brush border GO:0005903 Gene Ontology (GO) Relation: this pathophysiological event involves this cellular component This pathophysiological event involves Enterocyte brush border, annotated with brush border (GO:0005903). GO:0005903 is a cellular component from the Gene Ontology.
Small intestine UBERON:0002108 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Small intestine (UBERON:0002108). UBERON:0002108 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (2 references)
PMID:26358773 SUPPORT Human Clinical
"SLC9A3 encodes Na(+)/H(+) antiporter 3 (NHE3), which is the major intestinal brush-border Na(+)/H(+) exchanger."
Identifies the transporter whose loss defines the classic arm of this trigger and establishes it as the major brush-border sodium/proton exchanger.
PMID:26358773 SUPPORT Human Clinical
"This study identifies recessive mutations in NHE3, a downstream target of GC-C, as a cause of CSD"
States the relationship that makes this a single convergent node rather than two unrelated mechanisms: NHE3 is a downstream target of guanylate cyclase C, so activating GC-C and inactivating NHE3 act on the same axis.
Failure of Net Intestinal Sodium Absorption
The rate-limiting step. NHE3-mediated electroneutral sodium absorption is the principal route by which the small intestine and colon reclaim luminal sodium, and it is the driving force for the water absorption that follows it. When that step fails - whether the exchanger is absent, non-functional, or inhibited by excess cGMP - sodium is retained in the lumen at concentrations far above what the remaining absorptive routes can clear, and in the GUCY2C form active chloride secretion adds to the luminal ion load. This is the disorder-agnostic convergence point of all three genetic forms and the reason fecal sodium is disproportionately high, which is the diagnostic signature.
Small-intestinal enterocyte CL:0000584 Cell Ontology (CL) Relation: this pathophysiological event involves this cell type This pathophysiological event involves Small-intestinal enterocyte, annotated with enterocyte (CL:0000584). CL:0000584 is a cell type from the Cell Ontology.
Intestinal sodium absorption GO:0035725 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves decreased Intestinal sodium absorption, annotated with sodium ion transmembrane transport (GO:0035725). GO:0035725 is a biological process from the Gene Ontology. ↓ DECREASED Intestinal absorption GO:0050892 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves decreased Intestinal absorption (GO:0050892). GO:0050892 is a biological process from the Gene Ontology. ↓ DECREASED
Intestine UBERON:0000160 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Intestine (UBERON:0000160). UBERON:0000160 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (2 references)
PMID:19185281 SUPPORT Human Clinical
"Defective jejunal brush-border Na(+)/H(+) exchange has been reported in three sporadic patients"
Documents defective brush-border sodium/proton exchange measured directly in patient jejunum, the functional defect this node represents.
PMID:11113072 SUPPORT Human Clinical
"Congenital sodium diarrhea (CSD) is caused by defective sodium/proton exchange"
States the transport lesion underlying the disorder. Note that this paper's own negative conclusion - that CSD is unrelated to the known NHE genes including NHE3 - was later overturned by PMID:26358773, which identified recessive SLC9A3/NHE3 variants; only the sodium/proton-exchange defect claim is cited here.
Luminal Sodium Retention and Obligated Water Loss
Sodium retained in the lumen is osmotically active and obligates water, producing the high-volume watery stool. Because the retained solute is an ion the body itself secretes and recycles rather than an ingested nutrient, stool output does not fall when feeds are withheld and no dietary elimination abolishes it - the practical feature that separates this class from the diet-induced congenital diarrheas at the bedside. Diarrhea begins perinatally and, as in congenital chloride diarrhea, intrauterine fluid loss can present as polyhydramnios.
Transepithelial water movement into the lumen GO:0006833 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves increased Transepithelial water movement into the lumen, annotated with water transport (GO:0006833). GO:0006833 is a biological process from the Gene Ontology. ↑ INCREASED
Intestine UBERON:0000160 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Intestine (UBERON:0000160). UBERON:0000160 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"perinatal onset of a persistent watery diarrhea with nonproportionally high fecal sodium excretion"
States the output of this node - persistent watery stool carrying disproportionate sodium - and its perinatal onset.
Systemic Sodium Depletion and Metabolic Acidosis
The systemic arm, and the acid-base mirror image of congenital chloride diarrhea. NHE3 exports a proton into the lumen for every sodium it absorbs; when the exchanger fails, that proton is retained in the body while sodium is lost in stool. The result is hyponatremia with metabolic ACIDOSIS - the opposite derangement to the hypochloremic metabolic ALKALOSIS produced when the chloride/bicarbonate exchanger DRA fails in congenital chloride diarrhea. Urinary sodium is low-to-normal, confirming that the loss is enteric rather than renal, which is the discriminating measurement.
Systemic acid-base regulation GO:0006885 Gene Ontology (GO) Relation: this pathophysiological event involves this biological process This pathophysiological event involves abnormal Systemic acid-base regulation, annotated with regulation of pH (GO:0006885). GO:0006885 is a biological process from the Gene Ontology. ⚠ ABNORMAL
Show evidence (1 reference)
PMID:11113072 SUPPORT Human Clinical
"The diagnosis of CSD in 4 of 5 patients was based on daily fecal sodium excretion between 98 and 190 mmol/L, hyponatremia, metabolic acidosis, and low-to-normal urinary sodium concentrations."
Gives the complete biochemical signature of this node with quantitative fecal sodium values, including the low-to-normal urinary sodium that localizes the loss to the gut.
Electrolyte Crisis, Growth Failure and Later Enteral Adaptation
The clinical output. Mortality risk is high in the first years of life, from immediate electrolyte imbalance and from the complications of long-term parenteral nutrition. The natural history then diverges from that of the structural enteropathies in a clinically important way: survivors can eventually adapt to partial or complete enteral nutrition, so parenteral dependence is not necessarily lifelong and transplantation is not the expected endpoint. Management is supportive - aggressive sodium and fluid replacement with correction of acidosis - since no therapy addresses the transport defect itself.
Intestine UBERON:0000160 Uberon multi-species anatomy ontology (UBERON) Relation: this pathophysiological event occurs in this anatomical location This pathophysiological event occurs in Intestine (UBERON:0000160). UBERON:0000160 is an anatomical location from the Uberon multi-species anatomy ontology.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"Patients from both groups have a high risk of mortality due to immediate electrolyte imbalances and complications from long-term parenteral nutrition in the first years of life, but survivors can eventually adapt to partial or complete enteral nutrition."
States both halves of this node: the early mortality risk and the enteral adaptation achievable by survivors, which distinguishes the prognosis of CSD from that of the enterocyte-structural enteropathies.
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Pathograph

Use the checkboxes to hide or show graph categories. Hover nodes for evidence and cross-linked metadata.
Pathograph: causal mechanism network for Congenital Sodium Diarrhea Interactive directed graph showing how pathophysiology mechanisms, phenotypes, genetic factors and variants, experimental models, environmental triggers, and treatments relate through causal and linked edges.
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Phenotypes

7
Digestive 2
Watery Diarrhea with High Fecal Sodium OBLIGATE HP:0002014 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Watery diarrhea with high fecal sodium, annotated with Diarrhea (HP:0002014), qualified as temporality chronic. HP:0002014 is a phenotype from the Human Phenotype Ontology.
Temporal: CHRONIC
Show evidence (1 reference)
PMID:11113072 SUPPORT Human Clinical
"daily fecal sodium excretion between 98 and 190 mmol/L"
Quantifies the fecal sodium loss that defines the disorder.
Anal Atresia HP:0002023 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Anal atresia (HP:0002023). HP:0002023 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"CSD associated with choanal or anal atresia, hypertelorism, and corneal erosions--i.e., a syndromic form of CSD"
Names anal atresia among the defining features of the syndromic form.
Eye 2
Hypertelorism HP:0000316 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Hypertelorism (HP:0000316). HP:0000316 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"CSD associated with choanal or anal atresia, hypertelorism, and corneal erosions--i.e., a syndromic form of CSD"
Names hypertelorism among the defining features of the syndromic form.
Recurrent Corneal Erosions HP:0000495 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Recurrent corneal erosions (HP:0000495). HP:0000495 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"CSD associated with choanal or anal atresia, hypertelorism, and corneal erosions--i.e., a syndromic form of CSD"
Names corneal erosions among the defining features of the syndromic form.
Head and Neck 1
Choanal Atresia HP:0000453 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Choanal atresia (HP:0000453). HP:0000453 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"CSD associated with choanal or anal atresia, hypertelorism, and corneal erosions--i.e., a syndromic form of CSD"
Defines the syndromic form by this feature set, supporting the subtype assignment.
Metabolism 2
Metabolic Acidosis VERY_FREQUENT HP:0001942 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Metabolic acidosis (HP:0001942). HP:0001942 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:11113072 SUPPORT Human Clinical
"hyponatremia, metabolic acidosis, and low-to-normal urinary sodium concentrations"
Documents metabolic acidosis as part of the diagnostic biochemical triad. The VERY_FREQUENT band is a derived count: 4 of 5 patients in the cohort, i.e. 80%, the floor of the band. The cohort is a small sample of this entry's own population rather than a broader or narrower entity, so the band is scope-matched, but n=5 is thin and a larger series could move it.
Hyponatremia VERY_FREQUENT HP:0002902 Human Phenotype Ontology (HP) Relation: this clinical feature is this phenotype This clinical feature is Hyponatremia (HP:0002902). HP:0002902 is a phenotype from the Human Phenotype Ontology.
Show evidence (1 reference)
PMID:11113072 SUPPORT Human Clinical
"hyponatremia, metabolic acidosis, and low-to-normal urinary sodium concentrations"
Documents hyponatremia with low-to-normal urinary sodium in the diagnostic cohort. As for metabolic acidosis above, the VERY_FREQUENT band is the derived 4-of-5 count from this cohort; hyponatremia is also named in the disease definition, so the band is not resting on the count alone.
🧬

Genetic Associations

3
SLC9A3 loss-of-function variants (Causative)
Gene: SLC9A3 hgnc:11073 HUGO Gene Nomenclature Committee (hgnc) Relation: this disease-associated gene is this gene This disease-associated gene is SLC9A3 (hgnc:11073). hgnc:11073 is a gene from the HUGO Gene Nomenclature Committee. relationship_type: CAUSATIVE variant_origin: GERMLINE
Show evidence (2 references)
PMID:26358773 SUPPORT Human Clinical
"SLC9A3 missense, splicing and truncation mutations, including an instance of uniparental disomy, and whole-gene deletion were identified in nine patients from eight families with CSD."
The gene-discovery report identifying SLC9A3 as a cause of CSD and describing the variant spectrum.
PMID:26358773 SUPPORT In Vitro
"SLC9A3 missense mutations compromised NHE3 activity by reducing basal surface expression and/or loss of basal transport function of NHE3 molecules, whereas acute regulation was normal."
Functional characterization localizing the defect to constitutive NHE3 surface expression and transport rather than to acute regulation. Evidence source is IN_VITRO because activity was assayed in NHE-null fibroblasts.
GUCY2C activating variants (Causative)
Gene: GUCY2C hgnc:4688 HUGO Gene Nomenclature Committee (hgnc) Relation: this disease-associated gene is this gene This disease-associated gene is GUCY2C (hgnc:4688). hgnc:4688 is a gene from the HUGO Gene Nomenclature Committee. relationship_type: CAUSATIVE variant_origin: GERMLINE
Show evidence (2 references)
PMID:22436048 SUPPORT Human Clinical
"We identified a heterozygous missense mutation (c.2519G→T) in GUCY2C in all affected family members and observed no other rare variants in the exons of genes in the candidate region."
Identifies the causative heterozygous GUCY2C variant and its segregation with disease.
PMID:22436048 SUPPORT In Vitro
"Exposure of the mutant receptor to its ligands resulted in markedly increased production of cyclic guanosine monophosphate (cGMP)."
Demonstrates that the variant is activating, the point that distinguishes this form mechanistically from the loss-of-function forms. Evidence source is IN_VITRO because cGMP production was measured in transfected HEK293T cells.
SPINT2 loss-of-function variants (Causative)
Gene: SPINT2 hgnc:11247 HUGO Gene Nomenclature Committee (hgnc) Relation: this disease-associated gene is this gene This disease-associated gene is SPINT2 (hgnc:11247). hgnc:11247 is a gene from the HUGO Gene Nomenclature Committee. relationship_type: CAUSATIVE variant_origin: GERMLINE
Show evidence (2 references)
PMID:19185281 SUPPORT Human Clinical
"The same mutation and four distinct, homozygous or compound heterozygous mutations (p.Y163C, c.1A-->T, c.337+2T-->C, c.553+2T-->A) were identified in all syndromic patients. No SPINT2 mutations were found in classic-CSD patients."
Establishes SPINT2 as the syndromic-form gene and documents the clean genotype-phenotype separation from classic CSD.
PMID:19185281 SUPPORT In Vitro
"SPINT2 mutations were associated with loss of protein synthesis or failure to inhibit the serine protease trypsin in vitro."
Characterizes the two molecular consequences of the SPINT2 variants, supporting the excess-protease mechanism proposed for this form.
šŸ’Š

Medical Actions

1
Sodium and Fluid Replacement
Action: Nutritional SupportNCI Thesaurus (NCIT) Relation: this treatment is this clinical intervention This treatment is Nutritional Support (NCIT:C15433). NCIT:C15433 is a clinical intervention from the NCI Thesaurus. NCIT:C15433
Supportive management is the only available treatment: aggressive replacement of sodium and water with correction of the metabolic acidosis, delivered parenterally in infancy. Nothing addresses the underlying transport defect. Because survivors can adapt to partial or complete enteral nutrition, the goal is to carry the child through the high-mortality early years rather than to plan for lifelong parenteral support.
Mechanism Target:
INHIBITS Systemic Sodium Depletion and Metabolic Acidosis — Replacing sodium and water and correcting acidosis offsets the systemic consequences of enteric loss. It acts only on the systemic arm and leaves luminal sodium retention and stool volume unchanged.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"Patients from both groups have a high risk of mortality due to immediate electrolyte imbalances and complications from long-term parenteral nutrition in the first years of life"
Establishes electrolyte imbalance as the immediate threat to life that replacement therapy addresses, and records the competing risk from the parenteral route itself.
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Diagnosis

2
Fecal and urinary sodium measurement
Measurement of fecal sodium with paired serum electrolytes, blood gas and urinary sodium is the diagnostic core. The pattern is high fecal sodium with hyponatremia, metabolic acidosis and low-to-normal urinary sodium; the last of these localizes the loss to the gut rather than the kidney.
Show evidence (1 reference)
PMID:11113072 SUPPORT Human Clinical
"The diagnosis of CSD in 4 of 5 patients was based on daily fecal sodium excretion between 98 and 190 mmol/L, hyponatremia, metabolic acidosis, and low-to-normal urinary sodium concentrations."
States the diagnostic criteria actually used to establish CSD in a published cohort.
SLC9A3, GUCY2C and SPINT2 sequencing
Molecular confirmation. The presence or absence of atresias, hypertelorism and corneal erosions directs testing: syndromic features point to SPINT2, isolated intestinal disease to SLC9A3, and a dominant family history with inflammatory complications to GUCY2C.
Show evidence (1 reference)
PMID:19185281 SUPPORT Human Clinical
"No SPINT2 mutations were found in classic-CSD patients."
Supports using the syndromic/classic clinical split to direct molecular testing, since the genotype-phenotype separation is complete in the delineating cohort.
{ }

Source YAML

click to show
name: Congenital Sodium Diarrhea
creation_date: "2026-08-22T00:00:00Z"
category: Mendelian
synonyms:
- CSD
- congenital secretory sodium diarrhea
- diarrhea 8, secretory sodium, congenital
- familial GUCY2C diarrhea syndrome
description: >-
  Congenital sodium diarrhea (CSD) is a rare congenital diarrhea and enteropathy
  (CODE) defined by perinatal-onset watery diarrhea with disproportionately high
  fecal sodium loss, hyponatremia and metabolic ACIDOSIS. With congenital
  chloride diarrhea it forms the electrolyte-transport-related class of the CODEs
  - disorders in which the failed apical transport step handles an ion rather
  than an ingested nutrient, so the diarrhea is independent of feeding and cannot
  be treated by eliminating any dietary constituent. CSD is genetically
  heterogeneous but mechanistically convergent. Classic (non-syndromic) CSD is
  caused by recessive loss-of-function variants in SLC9A3, encoding NHE3, the
  major sodium/proton exchanger of the intestinal brush border. A second
  non-syndromic form is caused by dominant ACTIVATING variants in GUCY2C,
  encoding guanylate cyclase C: excess cyclic GMP both drives CFTR-mediated
  chloride and water secretion and inhibits NHE3, which is itself a downstream
  target of GC-C - so the gain-of-function and loss-of-function routes converge
  on the same failure of NHE3-mediated sodium absorption. A syndromic form is
  caused by recessive loss-of-function variants in SPINT2, encoding a Kunitz-type
  serine protease inhibitor, and is distinguished by choanal or anal atresia,
  hypertelorism and corneal erosions. CSD is the acid-base mirror image of
  congenital chloride diarrhea: losing the sodium/proton exchanger means protons
  are retained rather than secreted, giving acidosis where loss of the
  chloride/bicarbonate exchanger gives alkalosis. Mortality is high in the first
  years from electrolyte imbalance and the complications of parenteral nutrition,
  but survivors can adapt to partial or complete enteral nutrition - a natural
  history distinctly better than that of the structural enteropathies.
disease_term:
  preferred_term: congenital sodium diarrhea
  term:
    id: MONDO:0015170
    label: congenital sodium diarrhea
inheritance:
- name: Autosomal recessive inheritance
  description: >-
    Classic (SLC9A3) and syndromic (SPINT2) CSD are autosomal recessive.
    Consanguinity is common in reported kindreds; the original Austrian cohort
    traced five affected infants in a circumscribed rural area to a single common
    ancestor five generations back.
  inheritance_term:
    preferred_term: Autosomal recessive inheritance
    term:
      id: HP:0000007
      label: Autosomal recessive inheritance
  evidence:
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Pedigree analysis of the affected 2 CSD families revealed parental
      consanguinity and a common single ancestor 5 generations ago.
    explanation: >-
      Pedigree evidence for recessive inheritance with a founder effect in a
      geographically circumscribed population.
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Autosomal-recessive congenital sodium diarrhea (CSD) is characterized by
      perinatal onset of a persistent watery diarrhea with nonproportionally high
      fecal sodium excretion.
    explanation: >-
      States the recessive inheritance and the defining clinical and biochemical
      features of the disorder.
- name: Autosomal dominant inheritance
  description: >-
    The GUCY2C-associated form is autosomal dominant and results from activating
    (gain-of-function) rather than loss-of-function variants, so it does not
    follow the recessive pattern of the other two forms. It was delineated in a
    32-member Norwegian family. Applies to the GUCY2C-activating subtype only.
  inheritance_term:
    preferred_term: Autosomal dominant inheritance
    term:
      id: HP:0000006
      label: Autosomal dominant inheritance
  evidence:
  - reference: PMID:22436048
    reference_title: "Familial diarrhea syndrome caused by an activating GUCY2C mutation."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Familial diarrhea disorders are, in most cases, severe and caused by
      recessive mutations. We describe the cause of a novel dominant disease in
      32 members of a Norwegian family.
    explanation: >-
      Establishes the dominant inheritance of the GUCY2C form and explicitly
      contrasts it with the recessive majority of familial diarrheas.
genetic:
- name: SLC9A3 loss-of-function variants
  gene_term:
    preferred_term: SLC9A3
    term:
      id: hgnc:11073
      label: SLC9A3
  association: Causative
  relationship_type: CAUSATIVE
  variant_origin: GERMLINE
  subtype: Classic non-syndromic CSD
  features: >-
    Recessive missense, splicing, truncating and whole-gene-deletion variants in
    SLC9A3 (including one instance of uniparental disomy) cause classic
    non-syndromic CSD. All reported variants lie in the NHE3 N-terminal transport
    domain and all missense variants fall in putative membrane-spanning domains.
    Functional testing in NHE-null fibroblasts showed reduced basal surface
    expression and/or loss of basal transport function, with acute regulation
    preserved - so the defect is in constitutive NHE3 activity rather than in its
    regulation.
  evidence:
  - reference: PMID:26358773
    reference_title: "Reduced sodium/proton exchanger NHE3 activity causes congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      SLC9A3 missense, splicing and truncation mutations, including an instance
      of uniparental disomy, and whole-gene deletion were identified in nine
      patients from eight families with CSD.
    explanation: >-
      The gene-discovery report identifying SLC9A3 as a cause of CSD and
      describing the variant spectrum.
  - reference: PMID:26358773
    reference_title: "Reduced sodium/proton exchanger NHE3 activity causes congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      SLC9A3 missense mutations compromised NHE3 activity by reducing basal
      surface expression and/or loss of basal transport function of NHE3
      molecules, whereas acute regulation was normal.
    explanation: >-
      Functional characterization localizing the defect to constitutive NHE3
      surface expression and transport rather than to acute regulation. Evidence
      source is IN_VITRO because activity was assayed in NHE-null fibroblasts.
- name: GUCY2C activating variants
  gene_term:
    preferred_term: GUCY2C
    term:
      id: hgnc:4688
      label: GUCY2C
  association: Causative
  relationship_type: CAUSATIVE
  variant_origin: GERMLINE
  subtype: GUCY2C-activating form
  features: >-
    A heterozygous missense variant (c.2519G>T) in GUCY2C, encoding guanylate
    cyclase C, causes a dominant familial diarrhea syndrome that presents as
    non-syndromic CSD. The variant is activating, not inactivating: ligand
    exposure produces markedly increased cyclic GMP. Affected individuals have
    increased susceptibility to inflammatory bowel disease, small-bowel
    obstruction and esophagitis, an inflammatory dimension absent from the other
    forms.
  evidence:
  - reference: PMID:22436048
    reference_title: "Familial diarrhea syndrome caused by an activating GUCY2C mutation."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      We identified a heterozygous missense mutation (c.2519G→T) in GUCY2C in all
      affected family members and observed no other rare variants in the exons of
      genes in the candidate region.
    explanation: >-
      Identifies the causative heterozygous GUCY2C variant and its segregation
      with disease.
  - reference: PMID:22436048
    reference_title: "Familial diarrhea syndrome caused by an activating GUCY2C mutation."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      Exposure of the mutant receptor to its ligands resulted in markedly
      increased production of cyclic guanosine monophosphate (cGMP).
    explanation: >-
      Demonstrates that the variant is activating, the point that distinguishes
      this form mechanistically from the loss-of-function forms. Evidence source
      is IN_VITRO because cGMP production was measured in transfected HEK293T
      cells.
- name: SPINT2 loss-of-function variants
  gene_term:
    preferred_term: SPINT2
    term:
      id: hgnc:11247
      label: SPINT2
  association: Causative
  relationship_type: CAUSATIVE
  variant_origin: GERMLINE
  subtype: Syndromic CSD
  features: >-
    Homozygous or compound heterozygous SPINT2 variants, encoding a Kunitz-type
    serine protease inhibitor, cause the syndromic form. Reported alleles include
    a recurrent c.593-1G>A splice variant plus p.Y163C, c.1A>T, c.337+2T>C and
    c.553+2T>A. They act by loss of protein synthesis or by failure to inhibit
    the serine protease trypsin, so the proposed mechanism is excess serine
    protease activity in affected tissues. Genotype-phenotype separation is
    clean: SPINT2 variants were found in all syndromic patients and in none of
    the classic-CSD patients. SPINT2 variants also cause a syndromic form of
    congenital tufting enteropathy, so the same gene reaches two CODE classes.
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      The same mutation and four distinct, homozygous or compound heterozygous
      mutations (p.Y163C, c.1A-->T, c.337+2T-->C, c.553+2T-->A) were identified
      in all syndromic patients. No SPINT2 mutations were found in
      classic-CSD patients.
    explanation: >-
      Establishes SPINT2 as the syndromic-form gene and documents the clean
      genotype-phenotype separation from classic CSD.
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      SPINT2 mutations were associated with loss of protein synthesis or failure
      to inhibit the serine protease trypsin in vitro.
    explanation: >-
      Characterizes the two molecular consequences of the SPINT2 variants,
      supporting the excess-protease mechanism proposed for this form.
pathophysiology:
- name: Constitutive Guanylate Cyclase C Activation
  description: >-
    The upstream lesion of the GUCY2C arm only. A dominant activating variant in
    guanylate cyclase C raises cyclic GMP on ligand exposure. Excess cGMP acts in
    two directions: it hyperactivates CFTR, driving chloride and water secretion,
    and it inhibits NHE3, which is itself a downstream target of GC-C. This node
    is therefore placed upstream of the NHE3 lesion rather than beside it - the
    two are sequential in the GUCY2C arm, not alternative entry points at the
    same level. Conformers entering through SLC9A3 do not pass through this node.
  role: trigger
  biological_scale: MOLECULAR
  cell_types:
  - preferred_term: Small-intestinal enterocyte
    term:
      id: CL:0000584
      label: enterocyte
  molecular_functions:
  - preferred_term: Guanylate cyclase C activity
    term:
      id: GO:0004383
      label: guanylate cyclase activity
    modifier: GAIN_OF_FUNCTION
  biological_processes:
  - preferred_term: Cyclic GMP production
    term:
      id: GO:0006182
      label: cGMP biosynthetic process
    modifier: INCREASED
  locations:
  - preferred_term: Small intestine
    term:
      id: UBERON:0002108
      label: small intestine
  evidence:
  - reference: PMID:22436048
    reference_title: "Familial diarrhea syndrome caused by an activating GUCY2C mutation."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      Exposure of the mutant receptor to its ligands resulted in markedly
      increased production of cyclic guanosine monophosphate (cGMP).
    explanation: >-
      Demonstrates the gain-of-function that defines this node - markedly
      increased cGMP from the mutant receptor. Evidence source is IN_VITRO
      because cGMP was measured in transfected HEK293T cells.
  - reference: PMID:22436048
    reference_title: "Familial diarrhea syndrome caused by an activating GUCY2C mutation."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: >-
      This may cause hyperactivation of the cystic fibrosis transmembrane
      regulator (CFTR), leading to increased chloride and water secretion from
      the enterocytes, and may thus explain the chronic diarrhea in the affected
      family members.
    explanation: >-
      Gives the parallel secretory arm of this node. The authors hedge it as a
      proposed explanation, which is why the edge leaving this node is typed as
      having known intermediates rather than as direct.
  downstream:
  - target: Loss of NHE3-Mediated Apical Sodium/Proton Exchange
    causal_link_type: INDIRECT_KNOWN_INTERMEDIATES
- name: Loss of NHE3-Mediated Apical Sodium/Proton Exchange
  conforms_to: "electrolyte_transport_related_diarrhea#Loss of an Apical Intestinal Electrolyte Transport Step"
  description: >-
    The convergence point of every genetic form, and the node that carries the
    transport lesion itself. NHE3 is the major sodium/proton exchanger of the
    intestinal brush border. It is lost by two distinct routes that meet here
    rather than being alternatives at the same level: in classic CSD, recessive
    SLC9A3 variants directly reduce its surface expression or transport function
    while leaving acute regulation intact; in the GUCY2C form, excess cyclic GMP
    from the upstream activating variant inhibits it, NHE3 being a downstream
    target of GC-C. The syndromic SPINT2 form is proposed to act through excess
    serine protease activity in the affected epithelium, a route that is less
    well resolved at the transport level and is not modelled as a separate node
    here.
  role: trigger
  biological_scale: MOLECULAR
  cell_types:
  - preferred_term: Small-intestinal enterocyte
    term:
      id: CL:0000584
      label: enterocyte
  molecular_functions:
  - preferred_term: Brush-border sodium/proton exchange (NHE3)
    term:
      id: GO:0015385
      label: sodium:proton antiporter activity
    modifier: LOSS_OF_FUNCTION
  cellular_components:
  - preferred_term: Enterocyte brush border
    term:
      id: GO:0005903
      label: brush border
  locations:
  - preferred_term: Small intestine
    term:
      id: UBERON:0002108
      label: small intestine
  evidence:
  - reference: PMID:26358773
    reference_title: "Reduced sodium/proton exchanger NHE3 activity causes congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      SLC9A3 encodes Na(+)/H(+) antiporter 3 (NHE3), which is the major
      intestinal brush-border Na(+)/H(+) exchanger.
    explanation: >-
      Identifies the transporter whose loss defines the classic arm of this
      trigger and establishes it as the major brush-border sodium/proton
      exchanger.
  - reference: PMID:26358773
    reference_title: "Reduced sodium/proton exchanger NHE3 activity causes congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      This study identifies recessive mutations in NHE3, a downstream target of
      GC-C, as a cause of CSD
    explanation: >-
      States the relationship that makes this a single convergent node rather
      than two unrelated mechanisms: NHE3 is a downstream target of guanylate
      cyclase C, so activating GC-C and inactivating NHE3 act on the same axis.
  downstream:
  - target: Failure of Net Intestinal Sodium Absorption
    causal_link_type: DIRECT
- name: Failure of Net Intestinal Sodium Absorption
  conforms_to: "electrolyte_transport_related_diarrhea#Luminal Electrolyte Retention and Failure of Net Ion Absorption"
  description: >-
    The rate-limiting step. NHE3-mediated electroneutral sodium absorption is the
    principal route by which the small intestine and colon reclaim luminal
    sodium, and it is the driving force for the water absorption that follows it.
    When that step fails - whether the exchanger is absent, non-functional, or
    inhibited by excess cGMP - sodium is retained in the lumen at concentrations
    far above what the remaining absorptive routes can clear, and in the GUCY2C
    form active chloride secretion adds to the luminal ion load. This is the
    disorder-agnostic convergence point of all three genetic forms and the reason
    fecal sodium is disproportionately high, which is the diagnostic signature.
  role: central_effector
  biological_scale: TISSUE
  cell_types:
  - preferred_term: Small-intestinal enterocyte
    term:
      id: CL:0000584
      label: enterocyte
  biological_processes:
  - preferred_term: Intestinal sodium absorption
    term:
      id: GO:0035725
      label: sodium ion transmembrane transport
    modifier: DECREASED
  - preferred_term: Intestinal absorption
    term:
      id: GO:0050892
      label: intestinal absorption
    modifier: DECREASED
  locations:
  - preferred_term: Intestine
    term:
      id: UBERON:0000160
      label: intestine
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Defective jejunal brush-border Na(+)/H(+) exchange has been reported in
      three sporadic patients
    explanation: >-
      Documents defective brush-border sodium/proton exchange measured directly
      in patient jejunum, the functional defect this node represents.
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Congenital sodium diarrhea (CSD) is caused by defective sodium/proton
      exchange
    explanation: >-
      States the transport lesion underlying the disorder. Note that this
      paper's own negative conclusion - that CSD is unrelated to the known NHE
      genes including NHE3 - was later overturned by PMID:26358773, which
      identified recessive SLC9A3/NHE3 variants; only the sodium/proton-exchange
      defect claim is cited here.
  downstream:
  - target: Luminal Sodium Retention and Obligated Water Loss
    causal_link_type: DIRECT
  - target: Systemic Sodium Depletion and Metabolic Acidosis
    causal_link_type: DIRECT
- name: Luminal Sodium Retention and Obligated Water Loss
  conforms_to: "electrolyte_transport_related_diarrhea#Obligated Water Loss and Feeding-Independent Watery Diarrhea"
  description: >-
    Sodium retained in the lumen is osmotically active and obligates water,
    producing the high-volume watery stool. Because the retained solute is an ion
    the body itself secretes and recycles rather than an ingested nutrient, stool
    output does not fall when feeds are withheld and no dietary elimination
    abolishes it - the practical feature that separates this class from the
    diet-induced congenital diarrheas at the bedside. Diarrhea begins
    perinatally and, as in congenital chloride diarrhea, intrauterine fluid loss
    can present as polyhydramnios.
  role: effector
  biological_scale: TISSUE
  biological_processes:
  - preferred_term: Transepithelial water movement into the lumen
    term:
      id: GO:0006833
      label: water transport
    modifier: INCREASED
  locations:
  - preferred_term: Intestine
    term:
      id: UBERON:0000160
      label: intestine
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      perinatal onset of a persistent watery diarrhea with nonproportionally high
      fecal sodium excretion
    explanation: >-
      States the output of this node - persistent watery stool carrying
      disproportionate sodium - and its perinatal onset.
  downstream:
  - target: Electrolyte Crisis, Growth Failure and Later Enteral Adaptation
    causal_link_type: DIRECT
- name: Systemic Sodium Depletion and Metabolic Acidosis
  conforms_to: "electrolyte_transport_related_diarrhea#Ion-Specific Systemic Depletion and Acid-Base Derangement"
  description: >-
    The systemic arm, and the acid-base mirror image of congenital chloride
    diarrhea. NHE3 exports a proton into the lumen for every sodium it absorbs;
    when the exchanger fails, that proton is retained in the body while sodium is
    lost in stool. The result is hyponatremia with metabolic ACIDOSIS - the
    opposite derangement to the hypochloremic metabolic ALKALOSIS produced when
    the chloride/bicarbonate exchanger DRA fails in congenital chloride diarrhea.
    Urinary sodium is low-to-normal, confirming that the loss is enteric rather
    than renal, which is the discriminating measurement.
  role: effector
  biological_scale: ORGANISM
  biological_processes:
  - preferred_term: Systemic acid-base regulation
    term:
      id: GO:0006885
      label: regulation of pH
    modifier: ABNORMAL
  evidence:
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      The diagnosis of CSD in 4 of 5 patients was based on daily fecal sodium
      excretion between 98 and 190 mmol/L, hyponatremia, metabolic acidosis, and
      low-to-normal urinary sodium concentrations.
    explanation: >-
      Gives the complete biochemical signature of this node with quantitative
      fecal sodium values, including the low-to-normal urinary sodium that
      localizes the loss to the gut.
  downstream:
  - target: Electrolyte Crisis, Growth Failure and Later Enteral Adaptation
    causal_link_type: DIRECT
- name: Electrolyte Crisis, Growth Failure and Later Enteral Adaptation
  conforms_to: "electrolyte_transport_related_diarrhea#Electrolyte Crisis, Growth Failure and Long-Term Morbidity"
  description: >-
    The clinical output. Mortality risk is high in the first years of life, from
    immediate electrolyte imbalance and from the complications of long-term
    parenteral nutrition. The natural history then diverges from that of the
    structural enteropathies in a clinically important way: survivors can
    eventually adapt to partial or complete enteral nutrition, so parenteral
    dependence is not necessarily lifelong and transplantation is not the
    expected endpoint. Management is supportive - aggressive sodium and fluid
    replacement with correction of acidosis - since no therapy addresses the
    transport defect itself.
  role: consequence
  biological_scale: ORGANISM
  locations:
  - preferred_term: Intestine
    term:
      id: UBERON:0000160
      label: intestine
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Patients from both groups have a high risk of mortality due to immediate
      electrolyte imbalances and complications from long-term parenteral
      nutrition in the first years of life, but survivors can eventually adapt to
      partial or complete enteral nutrition.
    explanation: >-
      States both halves of this node: the early mortality risk and the enteral
      adaptation achievable by survivors, which distinguishes the prognosis of
      CSD from that of the enterocyte-structural enteropathies.
has_subtypes:
- name: Classic non-syndromic CSD
  display_name: Classic (non-syndromic) congenital sodium diarrhea, SLC9A3-related
  description: >-
    Recessive SLC9A3/NHE3 loss of function. Isolated intestinal disease with no
    atresias or facial features. Presented in seven families in the cohort that
    delineated the syndromic/classic split.
- name: Syndromic CSD
  display_name: Syndromic congenital sodium diarrhea, SPINT2-related
  description: >-
    Recessive SPINT2 loss of function. Intestinal disease plus choanal or anal
    atresia, hypertelorism and corneal erosions. Occurred in ten families in the
    delineating cohort. SPINT2 also causes a syndromic form of congenital
    tufting enteropathy.
- name: GUCY2C-activating form
  display_name: Familial GUCY2C diarrhea syndrome (dominant, activating)
  description: >-
    Dominant activating GUCY2C variant raising cyclic GMP. Milder, early-onset
    chronic diarrhea with increased susceptibility to inflammatory bowel disease,
    small-bowel obstruction and esophagitis. Paradoxically associated with
    prolonged rather than accelerated gut transit.
  genes:
  - preferred_term: GUCY2C
    term:
      id: hgnc:4688
      label: GUCY2C
  evidence:
  - reference: PMID:28957388
    reference_title: "Prolonged intestinal transit and diarrhea in patients with an activating GUCY2C mutation."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      Despite having diarrhea, the FGDS patients have prolonged transit time
      through the gut compared to HC, particularly in colon.
    explanation: >-
      Supports the paradoxical motility phenotype asserted for this subtype -
      diarrhea with prolonged rather than accelerated transit, most marked in
      the colon - which distinguishes it from the other CSD forms.
phenotypes:
- name: Watery Diarrhea with High Fecal Sodium
  category: Gastrointestinal
  description: >-
    Persistent watery diarrhea of perinatal onset carrying disproportionately
    high fecal sodium - between 98 and 190 mmol/L in the original cohort. It does
    not remit on withholding feeds.
  phenotype_term:
    preferred_term: Watery diarrhea with high fecal sodium
    term:
      id: HP:0002014
      label: Diarrhea
    temporality: CHRONIC
  frequency: OBLIGATE
  evidence:
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      daily fecal sodium excretion between 98 and 190 mmol/L
    explanation: >-
      Quantifies the fecal sodium loss that defines the disorder.
- name: Metabolic Acidosis
  category: Metabolic
  description: >-
    Metabolic acidosis from retention of protons that NHE3 would normally export
    into the lumen. This is the acid-base feature that distinguishes CSD from
    congenital chloride diarrhea, which causes metabolic alkalosis.
  phenotype_term:
    preferred_term: Metabolic acidosis
    term:
      id: HP:0001942
      label: Metabolic acidosis
  frequency: VERY_FREQUENT
  evidence:
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      hyponatremia, metabolic acidosis, and low-to-normal urinary sodium
      concentrations
    explanation: >-
      Documents metabolic acidosis as part of the diagnostic biochemical triad.
      The VERY_FREQUENT band is a derived count: 4 of 5 patients in the cohort,
      i.e. 80%, the floor of the band. The cohort is a small sample of this
      entry's own population rather than a broader or narrower entity, so the
      band is scope-matched, but n=5 is thin and a larger series could move it.
- name: Hyponatremia
  category: Metabolic
  description: >-
    Low serum sodium from continuous enteric sodium loss, with low-to-normal
    urinary sodium confirming the loss is not renal.
  phenotype_term:
    preferred_term: Hyponatremia
    term:
      id: HP:0002902
      label: Hyponatremia
  frequency: VERY_FREQUENT
  evidence:
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      hyponatremia, metabolic acidosis, and low-to-normal urinary sodium
      concentrations
    explanation: >-
      Documents hyponatremia with low-to-normal urinary sodium in the diagnostic
      cohort. As for metabolic acidosis above, the VERY_FREQUENT band is the
      derived 4-of-5 count from this cohort; hyponatremia is also named in the
      disease definition, so the band is not resting on the count alone.
- name: Choanal Atresia
  category: Craniofacial
  description: >-
    Choanal atresia occurs in the SPINT2-associated syndromic form and is one of
    the features that separates it from classic CSD.
  phenotype_term:
    preferred_term: Choanal atresia
    term:
      id: HP:0000453
      label: Choanal atresia
  subtype: Syndromic CSD
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      CSD associated with choanal or anal atresia, hypertelorism, and corneal
      erosions--i.e., a syndromic form of CSD
    explanation: >-
      Defines the syndromic form by this feature set, supporting the subtype
      assignment.
- name: Anal Atresia
  category: Gastrointestinal
  description: >-
    Anal atresia occurs as an alternative to choanal atresia in the syndromic
    form.
  phenotype_term:
    preferred_term: Anal atresia
    term:
      id: HP:0002023
      label: Anal atresia
  subtype: Syndromic CSD
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      CSD associated with choanal or anal atresia, hypertelorism, and corneal
      erosions--i.e., a syndromic form of CSD
    explanation: >-
      Names anal atresia among the defining features of the syndromic form.
- name: Hypertelorism
  category: Craniofacial
  description: >-
    Increased interocular distance, part of the facial phenotype of the
    SPINT2-associated syndromic form.
  phenotype_term:
    preferred_term: Hypertelorism
    term:
      id: HP:0000316
      label: Hypertelorism
  subtype: Syndromic CSD
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      CSD associated with choanal or anal atresia, hypertelorism, and corneal
      erosions--i.e., a syndromic form of CSD
    explanation: >-
      Names hypertelorism among the defining features of the syndromic form.
- name: Recurrent Corneal Erosions
  category: Ophthalmological
  description: >-
    Corneal erosions in the syndromic form, overlapping the ophthalmological
    phenotype seen in SPINT2-associated congenital tufting enteropathy.
  phenotype_term:
    preferred_term: Recurrent corneal erosions
    term:
      id: HP:0000495
      label: Recurrent corneal erosions
  subtype: Syndromic CSD
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      CSD associated with choanal or anal atresia, hypertelorism, and corneal
      erosions--i.e., a syndromic form of CSD
    explanation: >-
      Names corneal erosions among the defining features of the syndromic form.
diagnosis:
- name: Fecal and urinary sodium measurement
  description: >-
    Measurement of fecal sodium with paired serum electrolytes, blood gas and
    urinary sodium is the diagnostic core. The pattern is high fecal sodium with
    hyponatremia, metabolic acidosis and low-to-normal urinary sodium; the last
    of these localizes the loss to the gut rather than the kidney.
  evidence:
  - reference: PMID:11113072
    reference_title: "Congenital sodium diarrhea is an autosomal recessive disorder of sodium/proton exchange but unrelated to known candidate genes."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      The diagnosis of CSD in 4 of 5 patients was based on daily fecal sodium
      excretion between 98 and 190 mmol/L, hyponatremia, metabolic acidosis, and
      low-to-normal urinary sodium concentrations.
    explanation: >-
      States the diagnostic criteria actually used to establish CSD in a
      published cohort.
- name: SLC9A3, GUCY2C and SPINT2 sequencing
  description: >-
    Molecular confirmation. The presence or absence of atresias, hypertelorism
    and corneal erosions directs testing: syndromic features point to SPINT2,
    isolated intestinal disease to SLC9A3, and a dominant family history with
    inflammatory complications to GUCY2C.
  evidence:
  - reference: PMID:19185281
    reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: >-
      No SPINT2 mutations were found in classic-CSD patients.
    explanation: >-
      Supports using the syndromic/classic clinical split to direct molecular
      testing, since the genotype-phenotype separation is complete in the
      delineating cohort.
treatments:
- name: Sodium and Fluid Replacement
  description: >-
    Supportive management is the only available treatment: aggressive replacement
    of sodium and water with correction of the metabolic acidosis, delivered
    parenterally in infancy. Nothing addresses the underlying transport defect.
    Because survivors can adapt to partial or complete enteral nutrition, the
    goal is to carry the child through the high-mortality early years rather than
    to plan for lifelong parenteral support.
  therapeutic_modality: OTHER
  treatment_term:
    preferred_term: Nutritional Support
    term:
      id: NCIT:C15433
      label: Nutritional Support
  target_mechanisms:
  - target: Systemic Sodium Depletion and Metabolic Acidosis
    treatment_effect: INHIBITS
    description: >-
      Replacing sodium and water and correcting acidosis offsets the systemic
      consequences of enteric loss. It acts only on the systemic arm and leaves
      luminal sodium retention and stool volume unchanged.
    evidence:
    - reference: PMID:19185281
      reference_title: "Mutations in SPINT2 cause a syndromic form of congenital sodium diarrhea."
      supports: SUPPORT
      evidence_source: HUMAN_CLINICAL
      snippet: >-
        Patients from both groups have a high risk of mortality due to immediate
        electrolyte imbalances and complications from long-term parenteral
        nutrition in the first years of life
      explanation: >-
        Establishes electrolyte imbalance as the immediate threat to life that
        replacement therapy addresses, and records the competing risk from the
        parenteral route itself.
notes: >-
  CSD and congenital chloride diarrhea together form the
  electrolyte-transport-related class of the congenital diarrheas and
  enteropathies (PMID:29654747), and are curated as conformers of the
  electrolyte_transport_related_diarrhea module. Neither conforms to
  diet_induced_osmotic_diarrhea: the failed step handles an ion rather than an
  ingested nutrient, the diarrhea persists during bowel rest, and no dietary
  elimination abolishes it. The two are acid-base mirror images, which is the
  most useful bedside discriminator - alkalosis in chloride diarrhea, acidosis
  here.
  Curation caution on PMID:11113072: that paper's title asserts CSD is unrelated
  to the known sodium/proton exchanger genes, and it specifically excluded NHE3.
  That negative conclusion was overturned in 2015 by PMID:26358773, which
  identified recessive SLC9A3/NHE3 variants as a cause. The 2000 paper is cited
  here only for its inheritance, phenotype and biochemistry findings, which
  stand; it must not be cited for the claim that NHE3 is not involved.
  The GUCY2C form is curated here as a subtype rather than a separate entry
  because Janecke et al. classify it as a non-syndromic form of CSD, but it is
  atypical on three counts - dominant rather than recessive, gain- rather than
  loss-of-function, and accompanied by inflammatory bowel disease susceptibility.
  A future split into its own entry (familial GUCY2C diarrhea syndrome) would be
  defensible if its inflammatory arm is curated in depth.