A

Disease A

Slug:Alzheimer_Disease
B

Disease B

Cancer
Slug:Cancer
G

Causal Mechanism Graphs

Alzheimer Disease

graph LR
    Vascular_Dysfunction["Vascular Dysfunction"]
    Inhibitory_Neuron_Plasma_Proteomic_Age_Gap["Inhibitory Neuron Plasma Proteomic Age Gap"]
    Autophagy_Lysosomal_Dysfunction["Autophagy-Lysosomal Dysfunction"]
    Memory_Loss["Memory Loss"]
    PARP1_Mediated_Parthanatos["PARP1-Mediated Parthanatos"]
    Oligodendrocyte_Precursor_Cell_Plasma_Proteomic_Age_Gap["Oligodendrocyte Precursor Cell Plasma Proteomic Age Gap"]
    Necroptotic_Neuronal_Death["Necroptotic Neuronal Death"]
    PSEN1["PSEN1"]
    Complement_Mediated_Microglial_Synapse_Elimination["Complement-Mediated Microglial Synapse Elimination"]
    Tau_Induced_Nucleocytoplasmic_Transport_Failure["Tau-Induced Nucleocytoplasmic Transport Failure"]
    APOE["APOE"]
    Neurofibrillary_Tangle_Formation["Neurofibrillary Tangle Formation"]
    Tau_Directed_Antisense_Oligonucleotide_Therapy["Tau-Directed Antisense Oligonucleotide Therapy"]
    Neuroinflammation["Neuroinflammation"]
    Neuronal_Endosomal_Lysosomal_Acidification_Failure["Neuronal Endosomal-Lysosomal Acidification Failure"]
    APP["APP"]
    Amyloid_Plaque_Formation["Amyloid Plaque Formation"]
    Microglial_Lipid_Droplet_Accumulation["Microglial Lipid Droplet Accumulation"]
    Mitochondrial_Quality_Control_Failure["Mitochondrial Quality-Control Failure"]
    Glymphatic_Clearance_Failure["Glymphatic Clearance Failure"]
    Selective_Vulnerability_of_RORB+_Entorhinal_Excitatory_Neurons["Selective Vulnerability of RORB+ Entorhinal Excitatory Neurons"]
    Adaptive_Immune_T_Cell_Response_to_Tau_Pathology["Adaptive Immune T Cell Response to Tau Pathology"]
    Cumulative_exposure_to_strong_central_anticholinergic_medication["Cumulative exposure to strong central anticholinergic medication"]
    Oligodendrocyte_and_Myelin_Dysfunction["Oligodendrocyte and Myelin Dysfunction"]
    Interneuron_Dysfunction_and_Network_Hypersynchrony["Interneuron Dysfunction and Network Hypersynchrony"]
    LRP1["LRP1"]
    HSV_1_Reactivation_in_RORB+_Glutamatergic_Neurons["HSV-1 Reactivation in RORB+ Glutamatergic Neurons"]
    Synaptic_Dysfunction["Synaptic Dysfunction"]
    Muscarinic_M1_Receptor_Signaling_Loss["Muscarinic M1 Receptor Signaling Loss"]
    Oxidative_Stress["Oxidative Stress"]
    PSEN2["PSEN2"]
    Intercellular_Tau_Transmission_via_Extracellular_Vesicles["Intercellular Tau Transmission via Extracellular Vesicles"]
    Senescent_Cell_Accumulation["Senescent Cell Accumulation"]
    Astrocyte_Plasma_Proteomic_Age_Gap["Astrocyte Plasma Proteomic Age Gap"]

    Amyloid_Plaque_Formation --> Neurofibrillary_Tangle_Formation
    Amyloid_Plaque_Formation --> Synaptic_Dysfunction
    Amyloid_Plaque_Formation --> Neuroinflammation
    Neurofibrillary_Tangle_Formation --> Synaptic_Dysfunction
    Neurofibrillary_Tangle_Formation --> Necroptotic_Neuronal_Death
    Neurofibrillary_Tangle_Formation --> Intercellular_Tau_Transmission_via_Extracellular_Vesicles
    Synaptic_Dysfunction --> Memory_Loss
    Neuroinflammation --> Amyloid_Plaque_Formation
    Neuroinflammation --> Neurofibrillary_Tangle_Formation
    Neuroinflammation --> Oxidative_Stress
    Neuroinflammation --> Vascular_Dysfunction
    Oxidative_Stress --> Synaptic_Dysfunction
    Mitochondrial_Quality_Control_Failure --> Amyloid_Plaque_Formation
    Mitochondrial_Quality_Control_Failure --> Neurofibrillary_Tangle_Formation
    Mitochondrial_Quality_Control_Failure --> Memory_Loss
    Vascular_Dysfunction --> Amyloid_Plaque_Formation
    Vascular_Dysfunction --> Synaptic_Dysfunction
    Glymphatic_Clearance_Failure --> Amyloid_Plaque_Formation
    Autophagy_Lysosomal_Dysfunction --> Amyloid_Plaque_Formation
    Autophagy_Lysosomal_Dysfunction --> Neurofibrillary_Tangle_Formation
    HSV_1_Reactivation_in_RORB+_Glutamatergic_Neurons --> Synaptic_Dysfunction
    Intercellular_Tau_Transmission_via_Extracellular_Vesicles --> Neurofibrillary_Tangle_Formation
    PARP1_Mediated_Parthanatos --> Neuroinflammation
    PARP1_Mediated_Parthanatos --> Oxidative_Stress
    PARP1_Mediated_Parthanatos --> Mitochondrial_Quality_Control_Failure
    Adaptive_Immune_T_Cell_Response_to_Tau_Pathology --> Synaptic_Dysfunction
    Muscarinic_M1_Receptor_Signaling_Loss --> Amyloid_Plaque_Formation
    Muscarinic_M1_Receptor_Signaling_Loss --> Neurofibrillary_Tangle_Formation
    Complement_Mediated_Microglial_Synapse_Elimination --> Synaptic_Dysfunction
    Neuronal_Endosomal_Lysosomal_Acidification_Failure --> Amyloid_Plaque_Formation
    Neuronal_Endosomal_Lysosomal_Acidification_Failure --> Autophagy_Lysosomal_Dysfunction
    Selective_Vulnerability_of_RORB+_Entorhinal_Excitatory_Neurons --> Neurofibrillary_Tangle_Formation
    Necroptotic_Neuronal_Death --> Neuroinflammation
    Senescent_Cell_Accumulation --> Neurofibrillary_Tangle_Formation
    Senescent_Cell_Accumulation --> Neuroinflammation
    Oligodendrocyte_and_Myelin_Dysfunction --> Amyloid_Plaque_Formation
    Microglial_Lipid_Droplet_Accumulation --> Neurofibrillary_Tangle_Formation
    Microglial_Lipid_Droplet_Accumulation --> Neuroinflammation
    Interneuron_Dysfunction_and_Network_Hypersynchrony --> Memory_Loss
    Tau_Induced_Nucleocytoplasmic_Transport_Failure --> Neurofibrillary_Tangle_Formation
    Cumulative_exposure_to_strong_central_anticholinergic_medication --> Muscarinic_M1_Receptor_Signaling_Loss
    Tau_Directed_Antisense_Oligonucleotide_Therapy --> Neurofibrillary_Tangle_Formation
    Neuroinflammation -.-> Astrocyte_Plasma_Proteomic_Age_Gap
    Neurofibrillary_Tangle_Formation -.-> Astrocyte_Plasma_Proteomic_Age_Gap
    Neurofibrillary_Tangle_Formation -.-> Oligodendrocyte_Precursor_Cell_Plasma_Proteomic_Age_Gap
    Synaptic_Dysfunction -.-> Inhibitory_Neuron_Plasma_Proteomic_Age_Gap
    APP --> Amyloid_Plaque_Formation
    APP --> Neuronal_Endosomal_Lysosomal_Acidification_Failure
    APP --> Oligodendrocyte_and_Myelin_Dysfunction
    PSEN1 --> Amyloid_Plaque_Formation
    PSEN1 --> Neuronal_Endosomal_Lysosomal_Acidification_Failure
    PSEN2 --> Amyloid_Plaque_Formation
    APOE --> Microglial_Lipid_Droplet_Accumulation
    APOE --> Oligodendrocyte_and_Myelin_Dysfunction
    LRP1 --> Vascular_Dysfunction

    style Vascular_Dysfunction fill:#dbeafe
    style Inhibitory_Neuron_Plasma_Proteomic_Age_Gap fill:#e0e7ff
    style Autophagy_Lysosomal_Dysfunction fill:#dbeafe
    style Memory_Loss fill:#fef3c7
    style PARP1_Mediated_Parthanatos fill:#dbeafe
    style Oligodendrocyte_Precursor_Cell_Plasma_Proteomic_Age_Gap fill:#e0e7ff
    style Necroptotic_Neuronal_Death fill:#dbeafe
    style PSEN1 fill:#f3e8ff
    style Complement_Mediated_Microglial_Synapse_Elimination fill:#dbeafe
    style Tau_Induced_Nucleocytoplasmic_Transport_Failure fill:#dbeafe
    style APOE fill:#f3e8ff
    style Neurofibrillary_Tangle_Formation fill:#dbeafe
    style Tau_Directed_Antisense_Oligonucleotide_Therapy fill:#fce7f3
    style Neuroinflammation fill:#dbeafe
    style Neuronal_Endosomal_Lysosomal_Acidification_Failure fill:#dbeafe
    style APP fill:#f3e8ff
    style Amyloid_Plaque_Formation fill:#dbeafe
    style Microglial_Lipid_Droplet_Accumulation fill:#dbeafe
    style Mitochondrial_Quality_Control_Failure fill:#dbeafe
    style Glymphatic_Clearance_Failure fill:#dbeafe
    style Selective_Vulnerability_of_RORB+_Entorhinal_Excitatory_Neurons fill:#dbeafe
    style Adaptive_Immune_T_Cell_Response_to_Tau_Pathology fill:#dbeafe
    style Cumulative_exposure_to_strong_central_anticholinergic_medication fill:#dcfce7
    style Oligodendrocyte_and_Myelin_Dysfunction fill:#dbeafe
    style Interneuron_Dysfunction_and_Network_Hypersynchrony fill:#dbeafe
    style LRP1 fill:#f3e8ff
    style HSV_1_Reactivation_in_RORB+_Glutamatergic_Neurons fill:#dbeafe
    style Synaptic_Dysfunction fill:#dbeafe
    style Muscarinic_M1_Receptor_Signaling_Loss fill:#dbeafe
    style Oxidative_Stress fill:#dbeafe
    style PSEN2 fill:#f3e8ff
    style Intercellular_Tau_Transmission_via_Extracellular_Vesicles fill:#dbeafe
    style Senescent_Cell_Accumulation fill:#dbeafe
    style Astrocyte_Plasma_Proteomic_Age_Gap fill:#e0e7ff
H

Hypotheses

Tumour-suppressive amyloid/APP arm. Beyond the epidemiology, Alzheimer's disease-specific effectors provide a candidate mechanism for the protective direction: amyloid-beta (Abeta) and its precursor APP have tumour-suppressive and anti-angiogenic activity. Amyloid-beta oligomers inhibit the proliferation of human cancer cell lines in vitro, and transgenic mouse models of AD that overproduce Abeta show impaired growth and vascularization of implanted tumours - i.e., the AD-like, Abeta-rich brain environment is hostile to tumour growth. This is the AD-specific complement to the conserved aging/senescence tumour-suppressive barrier captured by the `senescence_tumor_suppression` module, and together they frame the cancer-AD inverse correlation as an antagonistic-pleiotropy trade-off in shared aging pathways rather than two independent phenomena.
PMID:31509551 (SUPPORT)
Source: IN_VITRO
"The effect of the cell growth inhibition, which was stronger in the case of HFIP Aβ oligomers, was observed for all cell lines."
In vitro evidence that amyloid-beta oligomers inhibit growth across multiple human cancer cell lines (leukemia, lung, breast), supporting a direct tumour-suppressive activity of an AD-defining molecule.
PMID:20739545 (SUPPORT)
Source: MODEL_ORGANISM
"Our data reveal that intracranial tumor growth and angiogenesis is significantly reduced in Tg APPsw and Tg PS1/APPsw mice compared with their wild-type littermates."
In vivo model-organism evidence that an Abeta-overproducing, AD-like brain environment suppresses orthotopic glioma growth and angiogenesis, supporting the protective direction at the tissue level. Model-organism evidence is kept distinct from the human epidemiological evidence above.
Shared molecular and inherited-genetic architecture. Independent of the AD-specific amyloid/APP arm, the inverse comorbidity has a candidate substrate in programs that are regulated in opposite directions between the AD and cancer states. Transcriptomic meta-analyses find many genes with inverse expression patterns in Alzheimer's disease versus lung cancer (and concordant patterns in Alzheimer's disease versus glioblastoma, a DIRECT comorbidity), with mitochondrial metabolism/oxidative phosphorylation a recurrent oppositely-regulated axis and the immune system a recurrent link. At the inherited level the signal is subtype-resolved rather than aggregate: genome-wide, Alzheimer's disease shows NO significant genetic correlation with cancer in the main (subtype-agnostic) datasets - reflecting little SHARED inherited risk rather than a negative correlation - and the inverse (negative) genetic signal emerges only on subtype resolution. Familial Alzheimer's disease shows strong NEGATIVE genetic correlations with lung cancer, and meta-analysis finds negative correlations with breast cancer, whereas late-onset Alzheimer's disease does not carry this negative signal - which is why subtype-agnostic estimates dilute the relationship. Mechanistically this converges on the amyloid arm of this entry: familial AD is precisely the subtype dominated by penetrant amyloid-processing biology (APP, PSEN1, PSEN2), the same effectors invoked in the tumour-suppressive amyloid/APP hypothesis above. The genetic-correlation evidence is from a non-peer-reviewed preprint (PPR:PPR1277612) and is recorded here as an emerging lead complementing the peer-reviewed transcriptomic and epidemiological evidence, not as sole support.
PMID:28667284 (SUPPORT)
Source: COMPUTATIONAL
"Transcriptomic meta-analyses reveal significant numbers of genes with inverse patterns of expression in Alzheimer's disease and lung cancer, and with similar patterns of expression in Alzheimer's disease and glioblastoma."
Peer-reviewed transcriptomic meta-analysis providing a molecular basis for the inverse (AD-lung cancer) and direct (AD-glioblastoma) comorbidities via oppositely- and concordantly-regulated gene sets.
PMID:28667284 (SUPPORT)
Source: COMPUTATIONAL
"Mitochondrial metabolism is regulated oppositely in Alzheimer's disease and lung cancer, indicating that it may be involved in the inverse co-morbidity between these diseases."
Identifies oppositely-regulated mitochondrial metabolism as a candidate molecular axis of the inverse comorbidity.
PPR:PPR1277612 (SUPPORT)
Source: COMPUTATIONAL
"No significant associations with cancer were detected for Alzheimer’s disease in the main-dataset analyses"
At the aggregate (subtype-agnostic) level the AD-cancer genetic correlation is null - little shared inherited risk, NOT a negative correlation - which is why the authors turned to subtype-resolved and meta-analytic approaches. Corrects any reading of low genetic affinity as itself an inverse signal. Non-peer-reviewed preprint; emerging lead.
PPR:PPR1277612 (SUPPORT)
Source: COMPUTATIONAL
"Familial Alzheimer’s disease showed strong negative correlations with lung cancer"
The subtype-resolved inverse genetic signal that actually supports this entry: familial AD is negatively genetically correlated with lung cancer (LDSC/HDL), whereas late-onset AD is not. Non-peer-reviewed preprint; emerging lead complementing the peer-reviewed evidence above.
PPR:PPR1277612 (SUPPORT)
Source: COMPUTATIONAL
"meta-analysis identified negative correlations with breast cancer in both methods"
Meta-analytic AD-cancer negative genetic correlation (breast cancer), concordant with the protective/inverse direction of this comorbidity. Non-peer-reviewed preprint; emerging lead, not sole support.
PPR:PPR1277612 (SUPPORT)
Source: COMPUTATIONAL
"familial Alzheimer’s disease is dominated by penetrant amyloid-processing biology involving APP , PSEN1 and PSEN2"
Mechanistic convergence with the "Tumour-suppressive amyloid/APP arm" hypothesis: the AD subtype carrying the negative (protective) genetic signal is the one driven by penetrant APP/PSEN amyloid biology, tying the inherited signal back to amyloid-beta/APP tumour suppression. Non-peer-reviewed preprint; emerging lead.
Y

Raw YAML

Show YAML
name: com_Alzheimer_Disease__Cancer
creation_date: "2026-07-03T14:04:34Z"
curation_status: CURATED
notes: >-
  The cancer/Alzheimer's-disease "inverse correlation" (or "cancer-AD paradox"):
  a history of cancer is associated with a reduced subsequent risk of Alzheimer's
  disease, and a diagnosis of Alzheimer's disease is associated with a reduced
  subsequent risk of cancer. This is a reciprocal (bidirectional) PROTECTIVE
  association, curated here as the flagship inverse comorbidity. Two features
  distinguish it from an artifact and from generic neurodegeneration. First, the
  major methodological biases (competing risk of death, diagnostic bias,
  selective survival) have been evaluated and are judged unlikely to fully
  explain the association (Ospina-Romero 2020), and the pattern is corroborated
  at the neuropathological level - a prior cancer diagnosis is associated with a
  lower burden of Alzheimer's-type neuropathology (Karanth 2022). Second, the
  inverse association appears relatively specific to Alzheimer's disease: it is
  seen for AD but not for vascular dementia (Roe 2010). Mechanistically this is
  framed as an evolutionary trade-off in which shared aging pathways are driven
  in opposite directions (cancer = sustained proliferation / growth-suppressor
  evasion / immune evasion; AD = excess neuronal death / growth-suppressor
  activation / immune activation), with AD-specific effectors such as APP and
  amyloid-beta acting as tumour suppressors. The antagonistic-pleiotropy
  substrate is captured by the `cellular_senescence` (deleterious arm) and
  `senescence_tumor_suppression` (tumour-suppressive arm) mechanism modules; the
  aging-associated loss of stemness node of the latter (PMID:39633048) is one
  conserved arm by which aging biology suppresses tumour initiation. Curation of
  this entry was prompted by the Comment "The cancer Alzheimer's disease paradox"
  (Feng, npj Aging 2026;12:93, doi:10.1038/s41514-026-00442-1). Note that the
  relationship is NOT uniformly protective across all neurodegeneration: cancer
  and Parkinson's disease show a more complex pattern (inverse for most cancers
  but a positive association with melanoma), which would be curated separately as
  effect_direction: MIXED. Two further nuances are captured in the "Shared
  molecular and inherited-genetic architecture" hypothesis: the AD-cancer
  relationship is inverse for lung cancer but DIRECT (positive) for glioblastoma
  (Sanchez-Valle 2017), and the inherited-genetic inverse signal is
  subtype-specific - familial AD carries a strong NEGATIVE (protective) genetic
  correlation with lung cancer while the aggregate, subtype-agnostic AD-cancer
  genetic correlation is null; late-onset AD does not carry the signal, so
  subtype-agnostic estimates dilute it. Familial AD's penetrant amyloid biology
  (APP/PSEN1/PSEN2) ties this back to the tumour-suppressive amyloid/APP arm.

disease_a:
  slug: Alzheimer_Disease
  preferred_term: Alzheimer disease
  term:
    id: MONDO:0004975
    label: Alzheimer disease

disease_b:
  slug: Cancer
  preferred_term: cancer
  term:
    id: MONDO:0004992
    label: cancer

directionality: BIDIRECTIONAL
effect_direction: PROTECTIVE

literature_evidence:
- reference: PMID:33185677
  reference_title: "Association Between Alzheimer Disease and Cancer With Evaluation of Study Biases: A Systematic Review and Meta-analysis."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "Observational studies consistently report inverse associations between cancer and Alzheimer disease (AD)."
  explanation: >-
    Systematic review and meta-analysis (22 studies, 9,630,435 individuals)
    establishing the consistently reported inverse (protective) association
    between cancer and Alzheimer's disease.
- reference: PMID:33185677
  reference_title: "Association Between Alzheimer Disease and Cancer With Evaluation of Study Biases: A Systematic Review and Meta-analysis."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "cancer was associated with decreased AD incidence (cohort studies: random-effects hazard ratio, 0.89; 95% CI, 0.79-1.00; case-control studies: random-effects odds ratio, 0.75; 95% CI, 0.61-0.93)."
  explanation: >-
    Pooled effect estimates below 1 quantify the protective direction: a prior
    cancer diagnosis is associated with lower Alzheimer's disease incidence.
- reference: PMID:33185677
  reference_title: "Association Between Alzheimer Disease and Cancer With Evaluation of Study Biases: A Systematic Review and Meta-analysis."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "Studies with insufficient or inappropriate confounder control or greater likelihood of AD diagnostic bias had mean hazard ratios closer to the null value, indicating that these biases could not explain the observed inverse association."
  explanation: >-
    Bias-adjusted metaregression argues the inverse association is not merely an
    artifact of confounding or diagnostic bias, supporting a genuine protective
    relationship.
- reference: PMID:22411920
  reference_title: "Inverse association between cancer and Alzheimer's disease: results from the Framingham Heart Study."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "Cancer survivors had a lower risk of Alzheimer's disease than those without cancer, and patients with Alzheimer's disease had a lower risk of incident cancer."
  explanation: >-
    Framingham Heart Study prospective cohort plus nested case-control analysis
    demonstrating the reciprocal (bidirectional) protective association in both
    directions.
- reference: PMID:20032288
  reference_title: "Cancer linked to Alzheimer disease but not vascular dementia."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "prevalent Alzheimer disease (AD) was longitudinally associated with a reduced risk of cancer, and a history of cancer was associated with a reduced risk of AD"
  explanation: >-
    Cardiovascular Health Study cohort independently confirming the reciprocal
    protective association between Alzheimer's disease and cancer.
- reference: PMID:20032288
  reference_title: "Cancer linked to Alzheimer disease but not vascular dementia."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "No significant association was found between cancer and subsequent development of VaD."
  explanation: >-
    Specificity evidence: the inverse association holds for Alzheimer's disease
    but not for vascular dementia, arguing the effect is AD-specific rather than
    a generic dementia/neurodegeneration phenomenon.
- reference: PMID:35094057
  reference_title: "Cancer diagnosis is associated with a lower burden of dementia and less Alzheimer's-type neuropathology."
  supports: SUPPORT
  evidence_source: HUMAN_CLINICAL
  snippet: "Cancer diagnosis was associated with a lower burden of Alzheimer's disease pathology and less cognitive impairment. These findings from a community-based cohort with neuropathological confirmation of substrates support the hypothesis that there is an inverse relationship between cancer and Alzheimer's disease."
  explanation: >-
    Autopsy cohort (University of Kentucky ADRC linked to the Kentucky Cancer
    Registry) providing neuropathological corroboration: prior cancer is
    associated with a lower burden of Alzheimer's-type pathology and cognitive
    impairment.

hypotheses:
- description: >-
    Tumour-suppressive amyloid/APP arm. Beyond the epidemiology, Alzheimer's
    disease-specific effectors provide a candidate mechanism for the protective
    direction: amyloid-beta (Abeta) and its precursor APP have tumour-suppressive
    and anti-angiogenic activity. Amyloid-beta oligomers inhibit the proliferation
    of human cancer cell lines in vitro, and transgenic mouse models of AD that
    overproduce Abeta show impaired growth and vascularization of implanted
    tumours - i.e., the AD-like, Abeta-rich brain environment is hostile to
    tumour growth. This is the AD-specific complement to the conserved
    aging/senescence tumour-suppressive barrier captured by the
    `senescence_tumor_suppression` module, and together they frame the cancer-AD
    inverse correlation as an antagonistic-pleiotropy trade-off in shared aging
    pathways rather than two independent phenomena.
  evidence:
  - reference: PMID:31509551
    reference_title: "Amyloid β oligomers inhibit growth of human cancer cells."
    supports: SUPPORT
    evidence_source: IN_VITRO
    snippet: "The effect of the cell growth inhibition, which was stronger in the case of HFIP Aβ oligomers, was observed for all cell lines."
    explanation: >-
      In vitro evidence that amyloid-beta oligomers inhibit growth across
      multiple human cancer cell lines (leukemia, lung, breast), supporting a
      direct tumour-suppressive activity of an AD-defining molecule.
  - reference: PMID:20739545
    reference_title: "Impaired orthotopic glioma growth and vascularization in transgenic mouse models of Alzheimer's disease."
    supports: SUPPORT
    evidence_source: MODEL_ORGANISM
    snippet: "Our data reveal that intracranial tumor growth and angiogenesis is significantly reduced in Tg APPsw and Tg PS1/APPsw mice compared with their wild-type littermates."
    explanation: >-
      In vivo model-organism evidence that an Abeta-overproducing, AD-like brain
      environment suppresses orthotopic glioma growth and angiogenesis,
      supporting the protective direction at the tissue level. Model-organism
      evidence is kept distinct from the human epidemiological evidence above.
- description: >-
    Shared molecular and inherited-genetic architecture. Independent of the
    AD-specific amyloid/APP arm, the inverse comorbidity has a candidate substrate
    in programs that are regulated in opposite directions between the AD and
    cancer states. Transcriptomic meta-analyses find many genes with inverse
    expression patterns in Alzheimer's disease versus lung cancer (and concordant
    patterns in Alzheimer's disease versus glioblastoma, a DIRECT comorbidity),
    with mitochondrial metabolism/oxidative phosphorylation a recurrent
    oppositely-regulated axis and the immune system a recurrent link. At the
    inherited level the signal is subtype-resolved rather than aggregate:
    genome-wide, Alzheimer's disease shows NO significant genetic correlation with
    cancer in the main (subtype-agnostic) datasets - reflecting little SHARED
    inherited risk rather than a negative correlation - and the inverse (negative)
    genetic signal emerges only on subtype resolution. Familial Alzheimer's
    disease shows strong NEGATIVE genetic correlations with lung cancer, and
    meta-analysis finds negative correlations with breast cancer, whereas
    late-onset Alzheimer's disease does not carry this negative signal - which is
    why subtype-agnostic estimates dilute the relationship. Mechanistically this
    converges on the amyloid arm of this entry: familial AD is precisely the
    subtype dominated by penetrant amyloid-processing biology (APP, PSEN1, PSEN2),
    the same effectors invoked in the tumour-suppressive amyloid/APP hypothesis
    above. The genetic-correlation evidence is from a non-peer-reviewed preprint
    (PPR:PPR1277612) and is recorded here as an emerging lead complementing the
    peer-reviewed transcriptomic and epidemiological evidence, not as sole support.
  evidence:
  - reference: PMID:28667284
    reference_title: "A molecular hypothesis to explain direct and inverse co-morbidities between Alzheimer's Disease, Glioblastoma and Lung cancer."
    supports: SUPPORT
    evidence_source: COMPUTATIONAL
    snippet: "Transcriptomic meta-analyses reveal significant numbers of genes with inverse patterns of expression in Alzheimer's disease and lung cancer, and with similar patterns of expression in Alzheimer's disease and glioblastoma."
    explanation: >-
      Peer-reviewed transcriptomic meta-analysis providing a molecular basis for
      the inverse (AD-lung cancer) and direct (AD-glioblastoma) comorbidities via
      oppositely- and concordantly-regulated gene sets.
  - reference: PMID:28667284
    reference_title: "A molecular hypothesis to explain direct and inverse co-morbidities between Alzheimer's Disease, Glioblastoma and Lung cancer."
    supports: SUPPORT
    evidence_source: COMPUTATIONAL
    snippet: "Mitochondrial metabolism is regulated oppositely in Alzheimer's disease and lung cancer, indicating that it may be involved in the inverse co-morbidity between these diseases."
    explanation: >-
      Identifies oppositely-regulated mitochondrial metabolism as a candidate
      molecular axis of the inverse comorbidity.
  - reference: PPR:PPR1277612
    reference_title: "A Genetic Atlas of Direct and Inverse Neuropsychiatric-Cancer Comorbidity"
    supports: SUPPORT
    evidence_source: COMPUTATIONAL
    snippet: "No significant associations with cancer were detected for Alzheimer’s disease in the main-dataset analyses"
    explanation: >-
      At the aggregate (subtype-agnostic) level the AD-cancer genetic correlation
      is null - little shared inherited risk, NOT a negative correlation - which is
      why the authors turned to subtype-resolved and meta-analytic approaches.
      Corrects any reading of low genetic affinity as itself an inverse signal.
      Non-peer-reviewed preprint; emerging lead.
  - reference: PPR:PPR1277612
    reference_title: "A Genetic Atlas of Direct and Inverse Neuropsychiatric-Cancer Comorbidity"
    supports: SUPPORT
    evidence_source: COMPUTATIONAL
    snippet: "Familial Alzheimer’s disease showed strong negative correlations with lung cancer"
    explanation: >-
      The subtype-resolved inverse genetic signal that actually supports this
      entry: familial AD is negatively genetically correlated with lung cancer
      (LDSC/HDL), whereas late-onset AD is not. Non-peer-reviewed preprint;
      emerging lead complementing the peer-reviewed evidence above.
  - reference: PPR:PPR1277612
    reference_title: "A Genetic Atlas of Direct and Inverse Neuropsychiatric-Cancer Comorbidity"
    supports: SUPPORT
    evidence_source: COMPUTATIONAL
    snippet: "meta-analysis identified negative correlations with breast cancer in both methods"
    explanation: >-
      Meta-analytic AD-cancer negative genetic correlation (breast cancer),
      concordant with the protective/inverse direction of this comorbidity.
      Non-peer-reviewed preprint; emerging lead, not sole support.
  - reference: PPR:PPR1277612
    reference_title: "A Genetic Atlas of Direct and Inverse Neuropsychiatric-Cancer Comorbidity"
    supports: SUPPORT
    evidence_source: COMPUTATIONAL
    snippet: "familial Alzheimer’s disease is dominated by penetrant amyloid-processing biology involving APP , PSEN1 and PSEN2"
    explanation: >-
      Mechanistic convergence with the "Tumour-suppressive amyloid/APP arm"
      hypothesis: the AD subtype carrying the negative (protective) genetic signal
      is the one driven by penetrant APP/PSEN amyloid biology, tying the inherited
      signal back to amyloid-beta/APP tumour suppression. Non-peer-reviewed
      preprint; emerging lead.
Source:GitHub