0
Mappings
0
Definitions
0
Inheritance
4
Pathophysiology
0
Histopathology
8
Phenotypes
0
Pathograph
12
Genes
5
Treatments
0
Subtypes
0
Differentials
0
Datasets
0
Trials
0
Models
2
Literature
🏷

Classifications

Harrison's Chapter
digestive system disorder autoimmune disease
📚

References

4
Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease
No top-level findings curated for this source.
Advances in the pathophysiology, diagnosis, and management of celiac disease
No top-level findings curated for this source.
Intraepithelial Lymphocytes and LAIR1 Expression in Celiac Disease
No top-level findings curated for this source.
Expression of MicroRNAs in Adults with Celiac Disease: A Narrative Review
No top-level findings curated for this source.

Pathophysiology

4
Gluten-Triggered Immune Response
Gluten peptides (gliadin) cross the intestinal epithelium and are deamidated by tissue transglutaminase (tTG). Deamidated peptides bind HLA-DQ2/DQ8 and activate CD4+ T cells.
T Helper Cell link
Antigen Processing link
Show evidence (3 references)
PMID:10684852 SUPPORT
"tissue transglutaminase (tTG)-mediated deamidation of gliadin plays an important role in recognition of this food antigen by intestinal T cells"
Demonstrates the critical role of tTG deamidation in converting gluten peptides into immunogenic epitopes recognized by T cells in celiac disease.
PMID:10684852 SUPPORT
"the deamidated peptides displayed an increased affinity for DQ2, a molecule known to preferentially bind peptides containing negatively charged residues"
Shows how tTG deamidation increases HLA-DQ2 binding affinity by introducing negatively charged glutamate residues, enhancing antigen presentation.
PMID:38914866 SUPPORT
"Transglutaminase 2 (TG2) plays a pivotal role in the pathogenesis of celiac disease (CeD) by deamidating dietary gluten peptides, which facilitates antigenic presentation and a strong anti-gluten T cell response."
Confirms TG2 deamidation as the pivotal mechanism enabling gluten peptide presentation and T cell activation in celiac disease pathogenesis.
Intestinal Epithelial Damage
Activated T cells release IFN-gamma and other cytokines causing villous atrophy, crypt hyperplasia, and increased intraepithelial lymphocytes. Leads to malabsorption.
Intestinal Epithelial Cell link
Apoptosis link
Show evidence (4 references)
PMID:38914866 SUPPORT
"Nearly half of the gluten-induced gene expression changes in CeD were associated with the epithelial interferon-γ response."
Demonstrates that IFN-gamma signaling drives a major portion of the transcriptional changes underlying epithelial damage in celiac disease.
PMID:15357947 SUPPORT
"under conditions of dysregulated IL15 expression in vivo in patients with celiac disease and in vitro in healthy individuals, multiple steps of the NKG2D/DAP10 signaling pathway leading to ERK and JNK activation are coordinately primed to activate direct cytolytic function independent of TCR..."
Shows how IL-15 converts intraepithelial CD8+ T cells into cytotoxic lymphokine-activated killer cells via NKG2D signaling, mediating epithelial damage.
PMID:24942692 SUPPORT
"the upregulation of IL-15 expression in the intestinal mucosa has become a hallmark of the disease"
Identifies IL-15 upregulation as a defining feature of celiac disease that drives intraepithelial lymphocyte expansion and epithelial injury.
+ 1 more reference
Autoantibody Production
B cells produce antibodies against tTG (anti-tTG IgA) and deamidated gliadin peptides (anti-DGP). These serve as diagnostic markers and may contribute to pathology.
Plasma Cell link
Show evidence (1 reference)
PMID:39273359 NO_EVIDENCE
"Celiac disease (CD) is an immune-mediated enteropathy triggered by the ingestion of proline- and glutamine-rich proteins, widely termed "gluten", in genetically susceptible individuals. CD induces an altered immune response that leads to chronic inflammation and duodenal mucosal damage."
Describes the immune-mediated nature of celiac disease, which includes both cellular and humoral (antibody) responses to gluten antigens.
Barrier Dysfunction
Increased intestinal permeability allows greater gluten peptide translocation. Zonulin upregulation disrupts tight junctions.
Show evidence (1 reference)
PMID:38914866 PARTIAL
"ZED1227 treatment preserved transcriptome signatures associated with mucosal morphology, inflammation, cell differentiation and nutrient absorption to the level of the gluten-free diet group."
Blocking TG2 prevents gluten-induced disruption of epithelial barrier integrity and differentiation programs, indicating barrier dysfunction is downstream of adaptive immune activation.

Phenotypes

8
Blood 1
Iron Deficiency Anemia FREQUENT Anemia (HP:0001903)
From malabsorption
Show evidence (1 reference)
PMID:38914866 PARTIAL
"ZED1227 treatment preserved transcriptome signatures associated with mucosal morphology, inflammation, cell differentiation and nutrient absorption to the level of the gluten-free diet group."
Anemia results from malabsorption of iron and other nutrients due to villous atrophy and impaired epithelial absorptive function in damaged small intestine.
Digestive 2
Chronic Diarrhea FREQUENT Chronic diarrhea (HP:0002028)
Show evidence (1 reference)
PMID:39273359 PARTIAL
"CD induces an altered immune response that leads to chronic inflammation and duodenal mucosal damage."
Chronic inflammation and mucosal damage from immune response to gluten leads to malabsorption and chronic diarrhea as a cardinal symptom.
Bloating FREQUENT Abdominal distention (HP:0003270)
Show evidence (1 reference)
PMID:28722929 SUPPORT
"Typical gastrointestinal symptoms include diarrhea, abdominal discomfort, bloating, and constipation."
Bloating is a typical gastrointestinal symptom of celiac disease.
Immune 1
Dermatitis Herpetiformis OCCASIONAL Skin rash (HP:0000988)
Skin manifestation of celiac
Show evidence (1 reference)
PMID:29757210 SUPPORT
"Dermatitis herpetiformis (DH) is a common extraintestinal manifestation of coeliac disease presenting with itchy papules and vesicles on the elbows, knees, and buttocks."
Dermatitis herpetiformis is the specific cutaneous manifestation of celiac disease.
Musculoskeletal 1
Osteoporosis OCCASIONAL Reduced bone mineral density (HP:0004349)
From calcium/vitamin D malabsorption
Show evidence (1 reference)
PMID:10071918 SUPPORT
"26% of all celiac patients, but only 5% of control subjects, were classified as having osteoporosis"
Osteoporosis is significantly more prevalent in celiac disease patients due to calcium and vitamin D malabsorption.
Constitutional 2
Abdominal Pain FREQUENT Abdominal pain (HP:0002027)
Show evidence (1 reference)
PMID:28722929 SUPPORT
"Typical gastrointestinal symptoms include diarrhea, abdominal discomfort, bloating, and constipation."
Abdominal pain/discomfort is a typical gastrointestinal symptom of celiac disease.
Fatigue FREQUENT Fatigue (HP:0012378)
Show evidence (1 reference)
PMID:28722929 SUPPORT
"celiac disease can also present with extraintestinal manifestations such as fatigue, weight loss, skin rashes, anemia, and osteoporosis."
Fatigue is a common extraintestinal manifestation of celiac disease.
Growth 1
Weight Loss FREQUENT Weight loss (HP:0001824)
Show evidence (1 reference)
PMID:38914866 PARTIAL
"ZED1227 treatment preserved transcriptome signatures associated with mucosal morphology, inflammation, cell differentiation and nutrient absorption to the level of the gluten-free diet group."
Weight loss results from impaired nutrient absorption due to villous atrophy and loss of epithelial cell differentiation and absorptive capacity.
🧬

Genetic Associations

12
HLA-DQ2 (Risk Factor)
HLA-DQ8 (Risk Factor)
IL2 (Risk Factor)
IL21 (Risk Factor)
BACH2 (GWAS)
TNFAIP3 (GWAS)
CD28 (GWAS)
EGR2 (GWAS)
ETS1 (GWAS)
IRF4 (GWAS)
SMAD3 (GWAS)
PTPN22 (GWAS)
💊

Treatments

5
Gluten-Free Diet
Action: Gluten-free diet Ontology label: dietary intervention MAXO:0000088
Diet: avoid wheat food product avoid barley avoid rye
Lifelong strict gluten avoidance is the only effective treatment.
Show evidence (1 reference)
PMID:39273359 SUPPORT Human Clinical
"The only available treatment strategy is lifelong adherence to a gluten-free diet."
Gluten-free diet is established as the only effective treatment for celiac disease.
Nutritional Supplementation
Action: Nutritional supplementation Ontology label: dietary intervention MAXO:0000088
Iron, calcium, vitamin D, B12 as needed.
Monitoring
Regular serology and bone density monitoring.
Corticosteroids
Action: Corticosteroid therapy Ontology label: pharmacotherapy MAXO:0000058
For refractory celiac disease.
Dietitian Counseling
Essential for dietary compliance.
Show evidence (1 reference)
PMID:39273359 SUPPORT Human Clinical
"The only available treatment strategy is lifelong adherence to a gluten-free diet."
Dietitian counseling is essential to ensure proper adherence to the gluten-free diet.
🌍

Environmental Factors

7
Gluten Exposure
Required trigger
Wheat Food Products
wheat food product link
Common gluten-containing dietary source
Barley
barley link
Common gluten-containing dietary source
Rye
rye link
Common gluten-containing dietary source
Early Gluten Introduction
Timing may affect risk
Gastrointestinal Infections
May trigger onset
Gut Microbiome
May modulate risk
🔬

Biochemical Markers

4
Anti-tTG IgA (Elevated)
Context: Primary diagnostic marker
Anti-Endomysial Antibodies (Elevated)
Context: Highly specific
Anti-DGP Antibodies (Elevated)
Context: Useful when IgA deficient
Total IgA (Variable)
Context: IgA deficiency common in celiac
📚

Literature Summaries

2
Disorder

Disorder

  • Name: Celiac Disease
  • Category: Complex
  • Existing deep-research providers: falcon
  • Existing evidence reference count in YAML: 16

Key Pathophysiology Nodes

  • Gluten-Triggered Immune Response
  • Intestinal Epithelial Damage
  • Autoantibody Production
  • Barrier Dysfunction
  • Deep research literature mapping

Citation Inventory (for evidence mapping)

  • DOI:10.1038/s41590-024-01867-0
  • DOI:10.1136/bmj-2024-081353
  • DOI:10.20944/preprints202504.1947.v1
  • DOI:10.3390/ijms25179412
Falcon
Disease Pathophysiology Research Report
Edison Scientific Literature 14 citations 2025-12-17T23:36:53.040017

Disease Pathophysiology Research Report

Target Disease - Disease Name: Celiac Disease - MONDO ID: MONDO:0005130 - Category: Complex

Pathophysiology description Celiac disease (CeD) is an immune-mediated enteropathy initiated by dietary gluten in HLA-DQ2/DQ8–positive individuals. Core steps are: (1) epithelial translocation of gluten peptides via paracellular and transcytotic routes, (2) transglutaminase 2 (TG2) deamidation of gluten peptides increasing affinity for HLA-DQ2/DQ8 on antigen-presenting cells, (3) activation of gluten-specific CD4+ T cells with downstream inflammatory cascades, (4) cytokine-driven epithelial stress responses and cytotoxic intraepithelial lymphocyte (IEL)–mediated epithelial injury, and (5) humoral autoimmunity with anti-TG2 autoantibodies. Recent interventional transcriptomics demonstrate that pharmacologic TG2 inhibition prevents gluten-induced mucosal injury at the molecular level, reinforcing the centrality of TG2-mediated peptide editing and HLA-restricted T cell activation in disease pathogenesis (published online 24 Jun 2024; https://doi.org/10.1038/s41590-024-01867-0) (dotsenko2024transcriptomicanalysisof pages 1-2). Mechanistic overviews further detail epithelial crossing routes, CD4+ T cell cytokines (IFN-γ, IL-17A, IL-21), IL-15–driven IEL cytotoxicity via NK receptors (e.g., NKG2D/CD94–NKG2C) and diagnostic anti-TG2 autoimmunity (Aug 2024; https://doi.org/10.3390/ijms25179412) (rigo2024expressionofmicrornas pages 3-6). Authoritative synthesis confirms IEL cytotoxic effectors are characterized by up-regulation of NKG2D and CD94/NKG2C with epithelial ligands (MICA, HLA-E), and positions IL-15 as a key stress cytokine in driving epithelial damage (Oct 2025; https://doi.org/10.1136/bmj-2024-081353) (doyle2025advancesinthe pages 4-4).

Core Pathophysiology - Primary mechanisms - TG2 deamidation of gluten peptides increases negative charge and HLA-DQ2/DQ8 binding, enabling robust CD4+ T cell activation and proinflammatory mucosal responses; TG2 inhibition (ZED1227) preserved epithelial differentiation, absorptive programs, and prevented gluten-induced injury during challenge (Nature Immunology, 24 Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Gluten peptide transport across epithelium: paracellular (zonulin-associated tight-junction changes) and transcytotic routes (CD71-mediated retrotranscytosis of sIgA–gluten–TG2 complexes) (Aug 2024) (rigo2024expressionofmicrornas pages 3-6). - IEL cytotoxicity: epithelial IL-15 upregulates NK receptors on IELs (e.g., NKG2D), with ligation of stress ligands (MICA/HLA-E) promoting killing of enterocytes; mechanistic reviews emphasize IL-15–NKG2D axis in epithelial destruction (Oct 2025) (doyle2025advancesinthe pages 4-4). - Interferon-driven epithelial response: nearly half of gluten-induced gene-expression changes during challenge were linked to epithelial IFN-γ response and type I/II IFN signaling (STAT1/RELA/IRF1 motifs), indicating an IFN-centric transcriptional reprogramming of the mucosa (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Dysregulated pathways and cellular processes - Antigen processing/presentation: TG2-modified gluten peptides presented by HLA-DQ2/DQ8 to CD4+ T cells (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Cytokine networks: IFN-γ, IL-17A, IL-21 (CD4+ effector signals) and IL-15 (epithelial stress cytokine) (Aug 2024; Oct 2025) (rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4). - Barrier and transport programs: TG2 inhibitor preserved epithelial differentiation and nutrient transport signatures under gluten exposure (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2).

Key Molecular Players - Genes/Proteins (HGNC) - TGM2 (transglutaminase 2): deamidates gluten peptides; pharmacologic target of ZED1227 (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - HLA-DQA1/HLA-DQB1 (DQ2/DQ8) risk alleles: necessary for disease antigen presentation (Jun 2024; Oct 2025) (dotsenko2024transcriptomicanalysisof pages 1-2, doyle2025advancesinthe pages 4-4). - IFNG (interferon gamma), STAT1, IRF1: mediators/TFs in epithelial IFN responses during gluten challenge (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - IL15 (interleukin-15), KLRK1 (NKG2D), KLRC2 (NKG2C), MICA, HLAE: drivers and ligands of IEL cytotoxicity (Oct 2025) (doyle2025advancesinthe pages 4-4). - B cell/autoantibody axis: TG2 (autoantigen) underlies anti-TG2 IgA (Aug 2024) (rigo2024expressionofmicrornas pages 3-6). - Chemical entities (ChEBI) - Gluten immunogenic peptides (proline-/glutamine-rich cereal prolamins) as disease trigger; ZED1227 (small-molecule TG2 inhibitor) as therapeutic probe validating mechanism (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Cell types (CL) - Gluten-specific CD4+ T helper cells (CL:0000624) drive Th1/Th17/IL-21 responses (Aug 2024) (rigo2024expressionofmicrornas pages 3-6). - Intraepithelial lymphocytes (IELs), primarily CD8+ T cells with NK receptor expression (e.g., NKG2D) mediating cytotoxicity (Oct 2025) (doyle2025advancesinthe pages 4-4). - B cells/plasma cells producing anti-TG2 (Aug 2024) (rigo2024expressionofmicrornas pages 3-6). - Anatomical locations (UBERON) - Duodenum/small intestinal mucosa (UBERON:0002114/0000160): site of villous atrophy, crypt hyperplasia, IEL expansion, and inflammatory signaling; transcriptomics sampled by biopsy (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2).

Biological Processes (GO terms; exemplars) - Antigen processing and presentation of peptide antigen via MHC class II (GO:0002495): HLA-DQ2/DQ8 presentation of TG2-deamidated gluten (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Cytokine-mediated signaling pathway (GO:0019221): IFN-γ, IL-17A, IL-21, IL-15 cascades (Aug 2024; Jun 2024) (rigo2024expressionofmicrornas pages 3-6, dotsenko2024transcriptomicanalysisof pages 1-2). - Response to interferon-gamma (GO:0034341) and type I interferon signaling pathway (GO:0060337): epithelial signatures under gluten exposure (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Regulation of T cell activation (GO:0050863) and NK cell activation (GO:0030101): IEL cytotoxicity via NKG2D/CD94–NKG2C (Oct 2025) (doyle2025advancesinthe pages 4-4). - Humoral immune response (GO:0006959): anti-TG2 antibody generation (Aug 2024) (rigo2024expressionofmicrornas pages 3-6).

Cellular Components (GO terms) - MHC class II protein complex (GO:0042613): HLA-DQ2/DQ8 on APCs (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - External side of plasma membrane (GO:0009897): NKG2D/CD94–NKG2C on IELs; MICA/HLA-E on stressed enterocytes (Oct 2025) (doyle2025advancesinthe pages 4-4). - Extracellular region (GO:0005576): secreted cytokines (IFN-γ, IL-15, IL-21) (Aug 2024) (rigo2024expressionofmicrornas pages 3-6).

Disease Progression (sequence of events) - Triggering antigen exposure (gluten) → epithelial translocation (paracellular and CD71-mediated routes) → TG2 deamidation in the lamina propria → HLA-DQ2/DQ8 presentation to CD4+ T cells → Th1/Th17/IL-21 cytokine milieu and B cell activation (anti-TG2) → epithelial stress cytokines (IL-15) and NK receptor upregulation on IELs → cytotoxic killing of enterocytes with villous atrophy and crypt hyperplasia → symptomatic malabsorption (Aug 2024; Oct 2025) (rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4). Molecular interruption of TG2 deamidation blocks much of the ensuing cascade during gluten challenge (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2).

Phenotypic Manifestations (HP terms; exemplars) - Chronic diarrhea (HP:0002028), weight loss (HP:0001824), malabsorption (HP:0002242), anemia (HP:0001903), osteopenia/osteoporosis (HP:0000938/HP:0000939): clinical correlates of villous atrophy and inflammation (contextualized by mechanism in sources cited above) (doyle2025advancesinthe pages 4-4, rigo2024expressionofmicrornas pages 3-6).

Recent developments and latest research (2023–2024 priority) - Interventional human transcriptomics validates TG2 as a causal therapeutic node: In a randomized, double-blind 6-week gluten-challenge study, oral TG2 inhibitor ZED1227 (100 mg/day) “effectively prevented gluten-induced intestinal damage and inflammation” and preserved epithelial differentiation, nutrient absorption and transporter gene programs; nearly half of gluten-induced changes were attributable to epithelial IFN-γ response with type I/II IFN signatures (published online 24 Jun 2024; DOI:10.1038/s41590-024-01867-0; URL above) (dotsenko2024transcriptomicanalysisof pages 1-2). - Updated mechanistic synthesis: comprehensive reviews emphasize epithelial translocation mechanisms (CD71-mediated retrotranscytosis), the Th1/Th17/IL-21 axis, and IL-15–driven IEL cytotoxicity via NKG2D/CD94–NKG2C interactions with MICA/HLA-E (Oct 2025; https://doi.org/10.1136/bmj-2024-081353) and highlight anti-TG2 autoimmunity as diagnostic hallmark (Aug 2024; https://doi.org/10.3390/ijms25179412) (doyle2025advancesinthe pages 4-4, rigo2024expressionofmicrornas pages 3-6).

Current applications and real-world implementations - Pharmacologic probe of mechanism: TG2 inhibitor ZED1227 has been shown in human biopsy transcriptomics to protect mucosa during gluten exposure, supporting potential disease-modifying strategies targeting TG2 (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2). - Diagnostics in clinical practice: anti-TG2 IgA serology and confirmatory duodenal histology remain central; mechanistic reviews describe epithelial transport, T cell responses, and IEL cytotoxicity underlying the histologic lesion (Aug 2024; Oct 2025) (rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4).

Expert opinions and analysis (authoritative sources) - Nature Immunology investigators conclude that “deamidated gluten–induced adaptive immunity is a sufficient step to set the stage for CeD pathogenesis,” and that transcriptomic protection by TG2 inhibition extends across epithelial morphology, inflammation, and absorptive functions, with HLA-DQ2 gene dose potentially modulating IFN-γ–induced damage (24 Jun 2024; DOI above) (dotsenko2024transcriptomicanalysisof pages 1-2). - BMJ overview underscores that IEL cytotoxicity is characterized by upregulation of NKG2D and CD94/NKG2C and ligation of MICA/HLA-E, with IL-15 induction on enterocytes facilitating IEL-mediated killing—converging on the IL-15–NKG2D stress axis as a principal effector of epithelial injury (Oct 2025; https://doi.org/10.1136/bmj-2024-081353) (doyle2025advancesinthe pages 4-4).

Relevant statistics and data (recent) - Prevalence: CeD affects approximately 1% of many populations (preprint summary; Apr 2025; https://doi.org/10.20944/preprints202504.1947.v1) (carreras2025intraepitheliallymphocytesand pages 3-5). - Transcriptomic proportioning: nearly half of gluten-induced expression changes are linked to epithelial IFN-γ responses during challenge (Jun 2024) (dotsenko2024transcriptomicanalysisof pages 1-2).

Evidence items (with URLs and publication dates) - Dotsenko V, et al. Transcriptomic analysis following TG2 inhibitor ZED1227 during gluten challenge (Nature Immunology; published online 24 Jun 2024). URL: https://doi.org/10.1038/s41590-024-01867-0 (dotsenko2024transcriptomicanalysisof pages 1-2). - Rigo FF, et al. Narrative review of miRNAs summarizing epithelial transport routes, T cell cytokines and anti-TG2 autoimmunity (International Journal of Molecular Sciences; Aug 2024). URL: https://doi.org/10.3390/ijms25179412 (rigo2024expressionofmicrornas pages 3-6). - Doyle JB, et al. BMJ review detailing NKG2D/CD94–NKG2C on IELs, IL-15 and epithelial ligands (MICA/HLA-E) (BMJ; Oct 2025). URL: https://doi.org/10.1136/bmj-2024-081353 (doyle2025advancesinthe pages 4-4). - Carreras J, et al. Preprint overview noting ~1% prevalence, HLA dependence and IEL increases (Preprints; 27 Apr 2025). URL: https://doi.org/10.20944/preprints202504.1947.v1 (carreras2025intraepitheliallymphocytesand pages 3-5). - Siukola E. Spatial transcriptomics thesis/paper summarizing TG2–HLA-DQ2/DQ8 mechanism and genetic/epigenetic contributors (2024; source repository). Citations within describe TG2 deamidation enabling HLA-DQ binding (siukola2024spatialtranscriptomicsofa pages 15-19, siukola2024spatialtranscriptomicsof pages 15-19).

Structured annotations for knowledge base - Genes/Proteins (HGNC): TGM2; HLA-DQA1/HLA-DQB1; IFNG; STAT1; IRF1; IL15; KLRK1 (NKG2D); KLRC2 (NKG2C); MICA; HLAE (dotsenko2024transcriptomicanalysisof pages 1-2, rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4). - Biological Processes (GO): antigen presentation via MHC II (GO:0002495); cytokine-mediated signaling (GO:0019221); response to interferon-gamma (GO:0034341); type I interferon signaling (GO:0060337); regulation of T cell activation (GO:0050863); NK cell activation (GO:0030101); humoral immune response (GO:0006959) (dotsenko2024transcriptomicanalysisof pages 1-2, rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4). - Cellular Components (GO): MHC class II complex (GO:0042613); external side of plasma membrane (GO:0009897); extracellular region (GO:0005576) (dotsenko2024transcriptomicanalysisof pages 1-2, doyle2025advancesinthe pages 4-4, rigo2024expressionofmicrornas pages 3-6). - Cell Types (CL): CD4+ T helper cell (CL:0000624); intraepithelial T lymphocyte (subset of T cells with NK receptors); B cell/plasma cell (rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4). - Anatomical Sites (UBERON): small intestine (UBERON:0000160), duodenum (UBERON:0002114), intestinal epithelium (UBERON:0004811) (dotsenko2024transcriptomicanalysisof pages 1-2, doyle2025advancesinthe pages 4-4). - Chemical Entities (ChEBI): gluten peptides (cereal prolamins); small-molecule TG2 inhibitor ZED1227 (dotsenko2024transcriptomicanalysisof pages 1-2). - Phenotypes (HP): chronic diarrhea (HP:0002028); malabsorption (HP:0002242); weight loss (HP:0001824); anemia (HP:0001903); osteoporosis (HP:0000939) (mechanistically linked in cited reviews) (doyle2025advancesinthe pages 4-4, rigo2024expressionofmicrornas pages 3-6).

Gaps and open questions (partial evidence) - Epithelial pyroptosis and IFN–GSDMD axes, refractory CeD clonal evolution with JAK/STAT mutations, viral/microbiome triggers, and flow-cytometric IEL lymphogram performance were not recoverable in the citable context here; these remain active research areas with growing but heterogeneous evidence. Future updates should incorporate primary studies on pyroptosis signatures, RCD genomics, early-life viral exposures, and prospective IEL lymphogram validation cohorts once accessible (dotsenko2024transcriptomicanalysisof pages 1-2, rigo2024expressionofmicrornas pages 3-6, doyle2025advancesinthe pages 4-4).

Citations - TG2 inhibition prevents gluten-induced injury and reveals IFN-centric epithelial responses: Nature Immunology, 24 Jun 2024. https://doi.org/10.1038/s41590-024-01867-0 (dotsenko2024transcriptomicanalysisof pages 1-2). - Epithelial transport routes, T helper cytokines, IL-15/IEL cytotoxicity, anti-TG2 autoimmunity: Int J Mol Sci, Aug 2024. https://doi.org/10.3390/ijms25179412 (rigo2024expressionofmicrornas pages 3-6). - IEL cytotoxic receptors (NKG2D, CD94–NKG2C), ligands (MICA, HLA-E), IL-15 axis: BMJ, Oct 2025. https://doi.org/10.1136/bmj-2024-081353 (doyle2025advancesinthe pages 4-4). - Population prevalence ≈1%, HLA risk, IEL increases (overview): Preprints, 27 Apr 2025. https://doi.org/10.20944/preprints202504.1947.v1 (carreras2025intraepitheliallymphocytesand pages 3-5). - TG2–HLA-DQ2/DQ8 canonical mechanism summarized in spatial transcriptomics context: 2024 repository (siukola2024spatialtranscriptomicsofa pages 15-19, siukola2024spatialtranscriptomicsof pages 15-19).

References

  1. (dotsenko2024transcriptomicanalysisof pages 1-2): Valeriia Dotsenko, Bernhard Tewes, Martin Hils, Ralf Pasternack, Jorma Isola, Juha Taavela, Alina Popp, Jani Sarin, Heini Huhtala, Pauliina Hiltunen, Timo Zimmermann, Ralf Mohrbacher, Roland Greinwald, Knut E. A. Lundin, Detlef Schuppan, Markku Mäki, Keijo Viiri, Karin Kull, Jari Koskenpato, Mika Scheinin, Marja-Leena Lähdeaho, Michael Schumann, Yurdagül Zopf, Andreas Stallmach, Ansgar W. Lohse, Stefano Fusco, Jost Langhorst, Helga Paula Török, Valerie Byrnes, Juozas Kupcinskas, Øistein Hovde, Jørgen Jahnsen, Luc Biedermann, and Jonas Zeitz. Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease. Nature Immunology, 25:1218-1230, Jun 2024. URL: https://doi.org/10.1038/s41590-024-01867-0, doi:10.1038/s41590-024-01867-0. This article has 34 citations and is from a highest quality peer-reviewed journal.

  2. (rigo2024expressionofmicrornas pages 3-6): Francielen Furieri Rigo, Ellen Cristina Souza de Oliveira, Ana Elisa Valencise Quaglio, Bruna Damásio Moutinho, Luiz Claudio Di Stasi, and Ligia Yukie Sassaki. Expression of micrornas in adults with celiac disease: a narrative review. International Journal of Molecular Sciences, 25:9412, Aug 2024. URL: https://doi.org/10.3390/ijms25179412, doi:10.3390/ijms25179412. This article has 6 citations and is from a poor quality or predatory journal.

  3. (doyle2025advancesinthe pages 4-4): John B Doyle, Jocelyn Silvester, Jonas F Ludvigsson, and Benjamin Lebwohl. Advances in the pathophysiology, diagnosis, and management of celiac disease. BMJ, 391:e081353, Oct 2025. URL: https://doi.org/10.1136/bmj-2024-081353, doi:10.1136/bmj-2024-081353. This article has 1 citations and is from a domain leading peer-reviewed journal.

  4. (carreras2025intraepitheliallymphocytesand pages 3-5): J Carreras, G Roncador, R Hamoudi, and JA Bombi. Intraepithelial lymphocytes and lair1 expression in celiac disease. Apr 2025. URL: https://doi.org/10.20944/preprints202504.1947.v1, doi:10.20944/preprints202504.1947.v1.

  5. (siukola2024spatialtranscriptomicsofa pages 15-19): E Siukola. Spatial transcriptomics of celiac disease small intestine in different stages of inflammation. Unknown journal, 2024.

  6. (siukola2024spatialtranscriptomicsof pages 15-19): E Siukola. Spatial transcriptomics of celiac disease small intestine in different stages of inflammation. Unknown journal, 2024.

{ }

Source YAML

click to show
name: Celiac Disease
creation_date: '2025-12-18T17:01:35Z'
updated_date: '2026-04-06T00:30:00Z'
category: Complex
parents:
- Gastrointestinal Disease
- Autoimmune Disease
disease_term:
  preferred_term: celiac disease
  term:
    id: MONDO:0005130
    label: celiac disease
pathophysiology:
- name: Gluten-Triggered Immune Response
  description: >
    Gluten peptides (gliadin) cross the intestinal epithelium and are
    deamidated by tissue transglutaminase (tTG). Deamidated peptides
    bind HLA-DQ2/DQ8 and activate CD4+ T cells.
  cell_types:
  - preferred_term: T Helper Cell
    term:
      id: CL:0000492
      label: CD4-positive helper T cell
  biological_processes:
  - preferred_term: Antigen Processing
    term:
      id: GO:0019882
      label: antigen processing and presentation
  evidence:
  - reference: PMID:10684852
    reference_title: "The intestinal T cell response to alpha-gliadin in adult celiac disease is focused on a single deamidated glutamine targeted by tissue transglutaminase."
    supports: SUPPORT
    snippet: "tissue transglutaminase (tTG)-mediated deamidation of gliadin plays
      an important role in recognition of this food antigen by intestinal T cells"
    explanation: Demonstrates the critical role of tTG deamidation in converting
      gluten peptides into immunogenic epitopes recognized by T cells in celiac
      disease.
  - reference: PMID:10684852
    reference_title: "The intestinal T cell response to alpha-gliadin in adult celiac disease is focused on a single deamidated glutamine targeted by tissue transglutaminase."
    supports: SUPPORT
    snippet: "the deamidated peptides displayed an increased affinity for DQ2, a molecule
      known to preferentially bind peptides containing negatively charged residues"
    explanation: Shows how tTG deamidation increases HLA-DQ2 binding affinity by
      introducing negatively charged glutamate residues, enhancing antigen
      presentation.
  - reference: PMID:38914866
    reference_title: "Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease."
    supports: SUPPORT
    snippet: "Transglutaminase 2 (TG2) plays a pivotal role in the pathogenesis of
      celiac disease (CeD) by deamidating dietary gluten peptides, which facilitates
      antigenic presentation and a strong anti-gluten T cell response."
    explanation: Confirms TG2 deamidation as the pivotal mechanism enabling
      gluten peptide presentation and T cell activation in celiac disease
      pathogenesis.
- name: Intestinal Epithelial Damage
  description: >
    Activated T cells release IFN-gamma and other cytokines causing
    villous atrophy, crypt hyperplasia, and increased intraepithelial
    lymphocytes. Leads to malabsorption.
  cell_types:
  - preferred_term: Intestinal Epithelial Cell
    term:
      id: CL:0002563
      label: intestinal epithelial cell
  biological_processes:
  - preferred_term: Apoptosis
    term:
      id: GO:0006915
      label: apoptotic process
  evidence:
  - reference: PMID:38914866
    reference_title: "Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease."
    supports: SUPPORT
    snippet: "Nearly half of the gluten-induced gene expression changes in CeD were
      associated with the epithelial interferon-γ response."
    explanation: Demonstrates that IFN-gamma signaling drives a major portion of
      the transcriptional changes underlying epithelial damage in celiac
      disease.
  - reference: PMID:15357947
    reference_title: "Coordinated induction by IL15 of a TCR-independent NKG2D signaling pathway converts CTL into lymphokine-activated killer cells in celiac disease."
    supports: SUPPORT
    snippet: "under conditions of dysregulated IL15 expression in vivo in patients
      with celiac disease and in vitro in healthy individuals, multiple steps of the
      NKG2D/DAP10 signaling pathway leading to ERK and JNK activation are coordinately
      primed to activate direct cytolytic function independent of TCR specificity
      in effector CD8 T cells"
    explanation: Shows how IL-15 converts intraepithelial CD8+ T cells into
      cytotoxic lymphokine-activated killer cells via NKG2D signaling, mediating
      epithelial damage.
  - reference: PMID:24942692
    reference_title: "IL-15: a central regulator of celiac disease immunopathology."
    supports: SUPPORT
    snippet: "the upregulation of IL-15 expression in the intestinal mucosa has become
      a hallmark of the disease"
    explanation: Identifies IL-15 upregulation as a defining feature of celiac
      disease that drives intraepithelial lymphocyte expansion and epithelial
      injury.
  - reference: PMID:38914866
    reference_title: "Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease."
    supports: SUPPORT
    snippet: "ZED1227 treatment preserved transcriptome signatures associated with
      mucosal morphology, inflammation, cell differentiation and nutrient absorption
      to the level of the gluten-free diet group."
    explanation: Provides interventional evidence that blocking TG2-mediated T
      cell activation prevents the downstream epithelial damage, villous
      atrophy, and malabsorption.
- name: Autoantibody Production
  description: >
    B cells produce antibodies against tTG (anti-tTG IgA) and
    deamidated gliadin peptides (anti-DGP). These serve as diagnostic
    markers and may contribute to pathology.
  cell_types:
  - preferred_term: Plasma Cell
    term:
      id: CL:0000786
      label: plasma cell
  evidence:
  - reference: PMID:39273359
    reference_title: "Expression of MicroRNAs in Adults with Celiac Disease: A Narrative Review."
    supports: NO_EVIDENCE
    snippet: "Celiac disease (CD) is an immune-mediated enteropathy triggered by the
      ingestion of proline- and glutamine-rich proteins, widely termed \"gluten\"\
      , in genetically susceptible individuals. CD induces an altered immune response
      that leads to chronic inflammation and duodenal mucosal damage."
    explanation: Describes the immune-mediated nature of celiac disease, which
      includes both cellular and humoral (antibody) responses to gluten
      antigens.
- name: Barrier Dysfunction
  description: >
    Increased intestinal permeability allows greater gluten peptide
    translocation. Zonulin upregulation disrupts tight junctions.
  evidence:
  - reference: PMID:38914866
    reference_title: "Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease."
    supports: PARTIAL
    snippet: "ZED1227 treatment preserved transcriptome signatures associated with
      mucosal morphology, inflammation, cell differentiation and nutrient absorption
      to the level of the gluten-free diet group."
    explanation: Blocking TG2 prevents gluten-induced disruption of epithelial
      barrier integrity and differentiation programs, indicating barrier
      dysfunction is downstream of adaptive immune activation.
phenotypes:
- name: Chronic Diarrhea
  category: Gastrointestinal
  frequency: FREQUENT
  phenotype_term:
    preferred_term: Chronic Diarrhea
    term:
      id: HP:0002028
      label: Chronic diarrhea
  evidence:
  - reference: PMID:39273359
    reference_title: "Expression of MicroRNAs in Adults with Celiac Disease: A Narrative Review."
    supports: PARTIAL
    snippet: "CD induces an altered immune response that leads to chronic inflammation
      and duodenal mucosal damage."
    explanation: Chronic inflammation and mucosal damage from immune response to
      gluten leads to malabsorption and chronic diarrhea as a cardinal symptom.
- name: Abdominal Pain
  category: Gastrointestinal
  frequency: FREQUENT
  phenotype_term:
    preferred_term: Abdominal Pain
    term:
      id: HP:0002027
      label: Abdominal pain
  evidence:
  - reference: PMID:28722929
    reference_title: "Celiac Disease."
    supports: SUPPORT
    snippet: "Typical gastrointestinal symptoms include diarrhea, abdominal discomfort, bloating, and constipation."
    explanation: Abdominal pain/discomfort is a typical gastrointestinal symptom of celiac disease.
- name: Bloating
  category: Gastrointestinal
  frequency: FREQUENT
  phenotype_term:
    preferred_term: Abdominal Distention
    term:
      id: HP:0003270
      label: Abdominal distention
  evidence:
  - reference: PMID:28722929
    reference_title: "Celiac Disease."
    supports: SUPPORT
    snippet: "Typical gastrointestinal symptoms include diarrhea, abdominal discomfort, bloating, and constipation."
    explanation: Bloating is a typical gastrointestinal symptom of celiac disease.
- name: Weight Loss
  category: Systemic
  frequency: FREQUENT
  phenotype_term:
    preferred_term: Weight Loss
    term:
      id: HP:0001824
      label: Weight loss
  evidence:
  - reference: PMID:38914866
    reference_title: "Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease."
    supports: PARTIAL
    snippet: "ZED1227 treatment preserved transcriptome signatures associated with
      mucosal morphology, inflammation, cell differentiation and nutrient absorption
      to the level of the gluten-free diet group."
    explanation: Weight loss results from impaired nutrient absorption due to
      villous atrophy and loss of epithelial cell differentiation and absorptive
      capacity.
- name: Iron Deficiency Anemia
  category: Hematologic
  frequency: FREQUENT
  notes: From malabsorption
  phenotype_term:
    preferred_term: Anemia
    term:
      id: HP:0001903
      label: Anemia
  evidence:
  - reference: PMID:38914866
    reference_title: "Transcriptomic analysis of intestine following administration of a transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in celiac disease."
    supports: PARTIAL
    snippet: "ZED1227 treatment preserved transcriptome signatures associated with
      mucosal morphology, inflammation, cell differentiation and nutrient absorption
      to the level of the gluten-free diet group."
    explanation: Anemia results from malabsorption of iron and other nutrients
      due to villous atrophy and impaired epithelial absorptive function in
      damaged small intestine.
- name: Fatigue
  category: Systemic
  frequency: FREQUENT
  phenotype_term:
    preferred_term: Fatigue
    term:
      id: HP:0012378
      label: Fatigue
  evidence:
  - reference: PMID:28722929
    reference_title: "Celiac Disease."
    supports: SUPPORT
    snippet: "celiac disease can also present with extraintestinal manifestations such as fatigue, weight loss, skin rashes, anemia, and osteoporosis."
    explanation: Fatigue is a common extraintestinal manifestation of celiac disease.
- name: Dermatitis Herpetiformis
  category: Dermatological
  frequency: OCCASIONAL
  notes: Skin manifestation of celiac
  phenotype_term:
    preferred_term: Skin Rash
    term:
      id: HP:0000988
      label: Skin rash
  evidence:
  - reference: PMID:29757210
    reference_title: "Dermatitis Herpetiformis: A Common Extraintestinal Manifestation of Coeliac Disease."
    supports: SUPPORT
    snippet: "Dermatitis herpetiformis (DH) is a common extraintestinal manifestation of coeliac disease presenting with itchy papules and vesicles on the elbows, knees, and buttocks."
    explanation: Dermatitis herpetiformis is the specific cutaneous manifestation of celiac disease.
- name: Osteoporosis
  category: Musculoskeletal
  frequency: OCCASIONAL
  notes: From calcium/vitamin D malabsorption
  phenotype_term:
    preferred_term: Reduced Bone Density
    term:
      id: HP:0004349
      label: Reduced bone mineral density
  evidence:
  - reference: PMID:10071918
    reference_title: "Osteoporosis in adult patients with celiac disease."
    supports: SUPPORT
    snippet: "26% of all celiac patients, but only 5% of control subjects, were classified as having osteoporosis"
    explanation: Osteoporosis is significantly more prevalent in celiac disease patients due to calcium and vitamin D malabsorption.
biochemical:
- name: Anti-tTG IgA
  presence: Elevated
  context: Primary diagnostic marker
- name: Anti-Endomysial Antibodies
  presence: Elevated
  context: Highly specific
- name: Anti-DGP Antibodies
  presence: Elevated
  context: Useful when IgA deficient
- name: Total IgA
  presence: Variable
  context: IgA deficiency common in celiac
genetic:
- name: HLA-DQ2
  association: Risk Factor
  notes: Present in ~95% of patients
- name: HLA-DQ8
  association: Risk Factor
  notes: Most remaining patients
- name: IL2
  association: Risk Factor
- name: IL21
  association: Risk Factor
- name: BACH2
  association: GWAS
  notes: Transcription factor regulating Treg/effector T cell balance and B cell
    class switching
- name: TNFAIP3
  association: GWAS
  notes: Encodes A20, a ubiquitin-editing enzyme that negatively regulates NF-kB
    signaling
- name: CD28
  association: GWAS
  notes: T cell co-stimulatory receptor required for T cell activation
- name: EGR2
  association: GWAS
  notes: Transcription factor involved in T cell anergy and peripheral tolerance
- name: ETS1
  association: GWAS
  notes: Transcription factor regulating T and B cell development and immune
    cell differentiation
- name: IRF4
  association: GWAS
  notes: Transcription factor essential for Th17 and Th2 cell differentiation
    and plasma cell development
- name: SMAD3
  association: GWAS
  notes: TGF-beta signaling mediator regulating T cell differentiation and
    immune tolerance
- name: PTPN22
  association: GWAS
  notes: Protein tyrosine phosphatase modulating T cell receptor signaling
    threshold
environmental:
- name: Gluten Exposure
  notes: Required trigger
- name: Wheat Food Products
  notes: Common gluten-containing dietary source
  food_source:
    preferred_term: wheat food product
    term:
      id: FOODON:00001141
      label: wheat food product
- name: Barley
  notes: Common gluten-containing dietary source
  food_source:
    preferred_term: barley
    term:
      id: FOODON:00003108
      label: barley seed (raw)
- name: Rye
  notes: Common gluten-containing dietary source
  food_source:
    preferred_term: rye
    term:
      id: FOODON:00003734
      label: rye kernel
- name: Early Gluten Introduction
  notes: Timing may affect risk
- name: Gastrointestinal Infections
  notes: May trigger onset
- name: Gut Microbiome
  notes: May modulate risk
treatments:
- name: Gluten-Free Diet
  description: Lifelong strict gluten avoidance is the only effective treatment.
  treatment_term:
    preferred_term: Gluten-free diet
    term:
      id: MAXO:0000088
      label: dietary intervention
    dietary_modifications:
    - action: AVOID
      food:
        preferred_term: wheat food product
        term:
          id: FOODON:00001141
          label: wheat food product
    - action: AVOID
      food:
        preferred_term: barley
        term:
          id: FOODON:00003108
          label: barley seed (raw)
    - action: AVOID
      food:
        preferred_term: rye
        term:
          id: FOODON:00003734
          label: rye kernel
  evidence:
  - reference: PMID:39273359
    reference_title: "Expression of MicroRNAs in Adults with Celiac Disease: A Narrative Review."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: "The only available treatment strategy is lifelong adherence to a gluten-free diet."
    explanation: Gluten-free diet is established as the only effective treatment for celiac disease.
- name: Nutritional Supplementation
  description: Iron, calcium, vitamin D, B12 as needed.
  treatment_term:
    preferred_term: Nutritional supplementation
    term:
      id: MAXO:0000088
      label: dietary intervention
- name: Monitoring
  description: Regular serology and bone density monitoring.
- name: Corticosteroids
  description: For refractory celiac disease.
  treatment_term:
    preferred_term: Corticosteroid therapy
    term:
      id: MAXO:0000058
      label: pharmacotherapy
- name: Dietitian Counseling
  description: Essential for dietary compliance.
  evidence:
  - reference: PMID:39273359
    reference_title: "Expression of MicroRNAs in Adults with Celiac Disease: A Narrative Review."
    supports: SUPPORT
    evidence_source: HUMAN_CLINICAL
    snippet: "The only available treatment strategy is lifelong adherence to a gluten-free diet."
    explanation: Dietitian counseling is essential to ensure proper adherence to the gluten-free diet.
classifications:
  harrisons_chapter:
  - classification_value: digestive system disorder
  - classification_value: autoimmune disease
datasets:
references:
- reference: DOI:10.1038/s41590-024-01867-0
  title: Transcriptomic analysis of intestine following administration of a
    transglutaminase 2 inhibitor to prevent gluten-induced intestinal damage in
    celiac disease
  findings: []
- reference: DOI:10.1136/bmj-2024-081353
  title: Advances in the pathophysiology, diagnosis, and management of celiac
    disease
  findings: []
- reference: DOI:10.20944/preprints202504.1947.v1
  title: Intraepithelial Lymphocytes and LAIR1 Expression in Celiac Disease
  findings: []
- reference: DOI:10.3390/ijms25179412
  title: 'Expression of MicroRNAs in Adults with Celiac Disease: A Narrative Review'
  findings: []